Non-enveloped, head-tail structure. The capsid is about 84-94 nm in diameter and has a T=16 icosahedral symmetry (Spounalikevirus) . The tail is 140-219 in length, contractile with globular structures at its tip, has 6 long terminal fibers, 6 short spikes and a double base plate.


Linear, dsDNA genome of 130-160 kb, encoding for about 190-230 proteins. Some Twortlikevirus might have circularly permuted genomes.


Genes are transcribed by operons.



  1. Adsorption: the phage attaches to the target cell through its tail fibers.
  2. Ejection of the viral DNA into host cell cytoplasm by contraction of the tail sheath.
  3. Transcription and translation of early genes.
  4. Replication of viral DNA. Transcription and translation of late genes.
  5. Assembly of an empty procapsid and packaging of the genome.
  6. Viral tail fibers assembly and viral tail assembly.
  7. Mature virions are released from the cell by lysis.

Matching UniProtKB/Swiss-Prot entries

48 entries grouped by strain (browse by keywords)

All proteins are shown (view only 13 complete proteome entries)

36 entries

Bacillus phage SP01 (Bacteriophage SP01) reference strain

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PORTL_BPSP1Probable portal protein
TSP_BPSP1Tail sheath protein (TSP)

8 entries

Enterococcus phage phiEF24C (Enterococcus bacteriophage phi-EF24C) reference strain

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BP40_BPPHEVirion protein 3 (Gene product 40) (gp40)
PORTL_BPPHEProbable portal protein
VPN1_BPPHEPutative tail sheath protein
VPN2_BPPHEPutative major capsid protein
VPN4_BPPHEVirion protein 4
VPN5_BPPHEVirion protein 5
VPN6_BPPHEVirion protein 6
VPN7_BPPHETail fiber protein

3 entries

Staphylococcus phage Twort (strain DSM 17442) (Bacteriophage Twort) reference strain

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GP168_BPTWOGene product 168 (gp168)
PORTL_BPTWOProbable portal protein
TSP_BPTWOTail sheath protein (TSP)

36 entries

Bacillus phage SP01 (Bacteriophage SP01)

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DPOL_BPSP1DNA polymerase (EC (EC 3.1.11.-)
DUHM_BPSP1Deoxyuridylate hydroxymethyltransferase (Deoxyuridylate hydroxymethylase) (EC ...
GP27_BPSP1Gene 27 protein
GP30_BPSP1Gene 30 protein (GP30)
GP33_BPSP1Gene 33 protein
GP37_BPSP1Putative gene 37 protein
GP38_BPSP1Putative gene 38 protein
GP39_BPSP1Putative gene 39 protein
GP40_BPSP1Gene 40 protein
GP41_BPSP1Putative gene 41 protein
GP42_BPSP1Putative gene 42 protein
GP43_BPSP1E22 protein (Gene 43 protein)
GP44_BPSP1E3 protein (Gene 44 protein)
GP45_BPSP1Putative gene 45 protein
GP46_BPSP1Putative gene 46 protein
GP47_BPSP1Putative gene 47 protein
GP48_BPSP1Putative gene 48 protein
GP49_BPSP1Putative gene 49 protein
GP50_BPSP1Putative gene 50 protein
GP51_BPSP1Putative gene 51 protein
GP52_BPSP1Putative gene 52 protein
GP53_BPSP1Putative gene 53 protein
GP54_BPSP1Putative gene 54 protein
GP55_BPSP1Putative gene 55 protein
GP56_BPSP1Putative gene 56 protein
GP57_BPSP1Putative gene 57 protein
GP58_BPSP1Putative gene 58 protein
GP59_BPSP1Putative gene 59 protein
GP60_BPSP1Putative gene 60 protein
HMUI_BPSP1DNA endonuclease I-HmuI (EC 3.1.-.-) (HNH homing endonuclease I-HmuI)
RP28_BPSP1RNA polymerase sigma GP28 factor
RP34_BPSP1RNA polymerase sigma GP34 factor
TF1_BPSP1Transcription factor 1
YG29_BPSP1Uncharacterized 16.2 kDa protein in gene 29 5'region

1 entry

Listeria phage A511 (Bacteriophage A511)

ALYS_BPA51Endolysin (EC (N-acetylmuramoyl-L-alanine amidase)