Enveloped, prolate ellipsoid form (ichnovirus) and cylindrical (bracovirus). Depending on the species, a virion contains one or several nucleocapsids.
Segmented, circular supercoiled dsDNAs ranging from 2.0-31kb. Each virion contains multiple copies of the genome.
a) : occurs in the calyx cells of the ovary.
- Excision and circularization of viral DNA in the ovary cells.
- Viral DNA synthesis.
- Appearance of viral envelopes and nucleocapsids in the nucleus.
- Mature virions are released into the lumen of the oviduct probably by lysis (brachovirus) or budding through the cell membrane (ichnovirus).
b) : occurs in all non-reproductive cells.
The viral genome is transmitted in proviral linear form integrated in the wasp's chromosomes. The virus DNA is replicated, just like any other part of the wasp's genome.