Partitiviridae (taxid:11012)



Non-enveloped icosahedral capsid, about 35-40 nm in diameter with a T=2* icosahedral symmetry . The two genomic segments are encapsidated separately.



Segmented linear dsRNA genome: 2 segments encoding potentially for 2 proteins. Segments are about 1.4-3.0 kb each, total size about 4 kb


Each genomic segment is monocistronic.




  1. Virus penetrates into the cytoplasm.
  2. Transcription of the dsRNA genome by viral polymerase occurs inside the virion, so that dsRNA is never exposed to the cytoplasm. This plus-strand transcript is used as template for translation.
  3. (+)RNAs are encapsidated in virion particle, inside which they are transcribed to give RNA (-) molecules with which they become base-paired to produce dsRNA genomes.
  4. Mature are transmitted to new cell by cytoplasmic exchange, sporogenesis or Hyphal anastomosis.

Matching UniProtKB/Swiss-Prot entries

(all links/actions below point to website)

6 entries grouped by protein

3 entries

Capsid protein (CP) (Coat protein)

Atkinsonella hypoxylon virus (isolate 2H) (AhV) reference strain
Cryptosporidium parvum virus 1 (strain KSU-1) reference strain
White clover cryptic virus 1 (isolate Boccardo/2004) (WCCV-1) reference strain

3 entries

RNA-directed RNA polymerase (EC

Atkinsonella hypoxylon virus (isolate 2H) (AhV) reference strain
Cryptosporidium parvum virus 1 (strain KSU-1) reference strain
White clover cryptic virus 1 (isolate Boccardo/2004) (WCCV-1) reference strain
These structures were created with the help of Colabfold by Jason Nomburg, Nathan Price and Jennifer Doudna (ModelArchive).

Amasya cherry disease-associated mycovirus taxid:284689

Protein ModelArchive
Putative coat protein ma-jd-viral-05199
RNA-dependent RNA polymerase ma-jd-viral-19149

Arabidopsis halleri partitivirus 1 taxid:1849335

Protein ModelArchive
Capsid protein ma-jd-viral-05179
RNA-dependent RNA polymerase ma-jd-viral-19147

Beet cryptic virus 1 taxid:509923

Protein ModelArchive
Coat protein ma-jd-viral-05194
Putative RNA-dependent RNA polymerase ma-jd-viral-19174

Cannabis cryptic virus taxid:1115692

Protein ModelArchive
Coat protein ma-jd-viral-09092
RNA-dependent RNA polymerase ma-jd-viral-19158

Ceratocystis resinifera virus 1 taxid:674982

Protein ModelArchive
Coat protein ma-jd-viral-09103
RNA-dependent RNA polymerase ma-jd-viral-19171

Dill cryptic virus 1 taxid:1408895

Protein ModelArchive
Coat protein ma-jd-viral-05198
RNA-dependent RNA polymerase ma-jd-viral-19162

Dill cryptic virus 2 taxid:1323529

Protein ModelArchive
Coat protein ma-jd-viral-09097
RNA-dependent RNA polymerase ma-jd-viral-19163

Discula destructiva virus 1 taxid:148880

Protein ModelArchive
RNA-dependent RNA polymerase ma-jd-viral-19812
Viral capsid protein ma-jd-viral-45144

Discula destructiva virus 2 taxid:160484

Protein ModelArchive
Capsid protein ma-jd-viral-45147
RNA-dependent RNA polymerase ma-jd-viral-19819

Fig cryptic virus taxid:882768

Protein ModelArchive
Putative capsid protein ma-jd-viral-34955
RNA-dependent RNA polymerase ma-jd-viral-19815

Fusarium poae partitivirus 2 taxid:1849533

Protein ModelArchive
Capsid protein ma-jd-viral-09087
RNA-dependent RNA polymerase ma-jd-viral-19144

Fusarium poae virus 1 taxid:75747

Protein ModelArchive
Putative capsid protein ma-jd-viral-09085

Fusarium poae virus 1-240374 taxid:1849534

Protein ModelArchive
Capsid protein ma-jd-viral-09086
RNA-dependent RNA polymerase ma-jd-viral-19141

Gremmeniella abietina RNA virus MS1 taxid:191436

Protein ModelArchive
Putative coat protein ma-jd-viral-45141
Putative replicase ma-jd-viral-19810
Uncharacterized protein ma-jd-viral-44341

Hop trefoil cryptic virus 2 taxid:1323523

Protein ModelArchive
Coat protein ma-jd-viral-09099
RNA-dependent RNA polymerase ma-jd-viral-19165

Penicillium aurantiogriseum partitivirus 1 taxid:1756157

Protein ModelArchive
47 kDa protein ma-jd-viral-45143
62 kDa protein ma-jd-viral-19820

Persimmon cryptic virus taxid:1183241

Protein ModelArchive
Coat protein ma-jd-viral-46659
RNA dependent RNA polymerase ma-jd-viral-19237

Pleurotus ostreatus virus 1 taxid:674983

Protein ModelArchive
Capsid protein ma-jd-viral-09090
RNA-dependent RNA polymerase ma-jd-viral-19173

Primula malacoides virus China/Mar2007 taxid:479713

Protein ModelArchive
Putative RNA-dependent RNA polymerase ma-jd-viral-19157
Putative capsid protein ma-jd-viral-09094

Pseudogymnoascus destructans partitivirus-pa taxid:1685502

Protein ModelArchive
Capsid protein ma-jd-viral-45146
RNA-dependent RNA polymerase ma-jd-viral-19821

Raphanus sativus cryptic virus 1 taxid:348449

Protein ModelArchive
Coat protein ma-jd-viral-05188
RNA-dependent RNA polymerase ma-jd-viral-19180

Raphanus sativus cryptic virus 2 taxid:351495

Protein ModelArchive
Coat protein ma-jd-viral-56145
Coat protein ma-jd-viral-56146
Putative RNA dependent RNA polymerse ma-jd-viral-19813

Raphanus sativus cryptic virus 3 taxid:580254

Protein ModelArchive
Coat protein ma-jd-viral-46658
Putative RNA-dependent RNA polymerase ma-jd-viral-19822

Red clover cryptic virus 1 taxid:1408894

Protein ModelArchive
Coat protein ma-jd-viral-05190
RNA-dependent RNA polymerase ma-jd-viral-19168

Red clover cryptic virus 2 taxid:1323524

Protein ModelArchive
Coat protein ma-jd-viral-09096
RNA-dependent RNA polymerase ma-jd-viral-19177

Rhizoctonia solani dsRNA virus 2 taxid:1411681

Protein ModelArchive
Coat protein ma-jd-viral-05193
RNA-dependent RNA polymerase ma-jd-viral-19181

Rhizoctonia solani dsRNA virus 3 taxid:1825688

Protein ModelArchive
RNA-dependent RNA polymerase ma-jd-viral-19153

Rosellinia necatrix partitivirus 1-W8 taxid:235994

Protein ModelArchive
Putative RNA-dependent RNA polymerase ma-jd-viral-19166
Putative coat protein ma-jd-viral-09093

Rosellinia necatrix partitivirus 2 taxid:859651

Protein ModelArchive
Capsid protein ma-jd-viral-05180
RNA-dependent RNA polymerase ma-jd-viral-19143

Rosellinia necatrix partitivirus 6 taxid:1573459

Protein ModelArchive
Capsid protein ma-jd-viral-09095
RNA dependent RNA polymerase ma-jd-viral-19164

Sclerotinia sclerotiorum partitivirus S taxid:659497

Protein ModelArchive
Putative coat protein ma-jd-viral-05189
RNA-dependent RNA polymerase ma-jd-viral-19155

Sinapis alba cryptic virus 1 taxid:1768874

Protein ModelArchive
Capsid protein ma-jd-viral-46652
RNA dependent RNA polymerase ma-jd-viral-19807

Sophora japonica powdery mildew-associated partitivirus taxid:1891704

Protein ModelArchive
Putative RNA-dependent RNA polymerase ma-jd-viral-19167
Putative coat protein ma-jd-viral-05187

Spinach cryptic virus 1 taxid:1937817

Protein ModelArchive
Coat protein ma-jd-viral-05201
RNA-dependent RNA polymerase ma-jd-viral-19175

Ustilaginoidea virens partitivirus 2 taxid:1415665

Protein ModelArchive
Coat protein ma-jd-viral-32253
RNA-dependent RNA polymerase ma-jd-viral-19814

Verticillium dahliae partitivirus 1 taxid:1310424

Protein ModelArchive
Coat protein ma-jd-viral-45145
RNA-dependent RNA polymerase ma-jd-viral-19818

Vicia cryptic virus taxid:54289

Protein ModelArchive
Coat protein ma-jd-viral-05192
Putative RNA dependent RNA polymerase ma-jd-viral-19154

White clover cryptic virus 2 taxid:1323525

Protein ModelArchive
Coat protein ma-jd-viral-09091
RNA-dependent RNA polymerase ma-jd-viral-19160