Non-enveloped, head-tail structure. The icosahedral head is about 85 nm in diameter and virions also produce head size variants (about 47-65 nm). The tail is long, 228 × 18 nm, contractile, has six 90 nm-long kinked fibers and a base plate.
Linear, dsDNA genome of about 100 kb, encoding for approximately 100 proteins. The genome is terminally redundant and circularly permuted.
Genes are transcribed by operons.
- Adsorption: the phage attaches to the target cell through its tail fibers. Two types of fibers are encoded by the phage to allow receptor tropism switching.
- Ejection of the viral DNA into host cell cytoplasm by contraction of the tail sheath.
- Transcription and translation of early genes.
- Synthesis of linear concatemers copies of viral DNA. Transcription and translation of late genes.
- Assembly of an empty procapsid and packaging of the genome.
- Viral tail fibers assembly and viral tail assembly.
- Mature virions are released from the cell by lysis.