Ribosomal frameshifting

Programmed Ribosomal frameshifting is an alternate mechanism of translation to merge proteins encoded by two overlapping open reading frames. The frameshift occurs at low frequency and consists of ribosomes slipping by one base in either the 5'(-1) or 3'(+1) directions during translation. Some viruses contains both a +1 and a -1 ribosomal frameshifts

-1 Ribosomal frameshift

All cis-acting frameshift signals encoded in mRNAs are minimally composed of two functional elements: a heptanucleotide “slippery sequence” conforming to the general form X XXY YYZ, followed by a RNA structural element, usually a H-type RNA pseudoknot, positioned an optimal number of nucleotides (5 to 9) downstream. Mechanistically, the ribosome is stalled on the slippery sequence by the pseudoknot structure in 3'. In about 10% cases, the ribosome will backtrack one nucleotide, creating one or more mismatchs between tRNAs and mRNA. Translation resolves on the backtracked ribosome in the -1 frame.

Family Genus Virus Type RefSeq .......... XXXYYY Z .......... Frameshift position Freq Ref
-1/-2 FRAMESHIFTS
Totiviridae Giardiavirus Giardia lamblia virus -1 NC_003555 CGUGCGCCAU CCCUUU A UCCGAUCGUG 2871
Totivirus Saccharomyces cerevisiae virus L-A -1 NC_003745 GUACUCAGCA GGGUUU A GGAGUGGUAG 1964
Astroviridae Avastrovirus Turkey astrovirus 1 -1 NC_002470 CUACGUGUUC AAAAAA C UAGAUAGUCA 3307
Mamastrovirus Human astrovirus 1 -1 NC_001943 ACAAGGCCCC AAAAAA C UACAAAGGGC 2845
Luteoviridae Enamovirus Pea enation mosaic virus-1 -1 NC_003629 CCAGACGCUC GGGAAA C GGAUUAUUCC 2035
Luteovirus Barley yellow dwarf virus-PAV -1 NC_004750 TTGACTCTGT GGGUUU U AGAGGGGCTC 1159
Polerovirus Potato leafroll virus -1 NC_001747 CAAACAAGCC UUUAAA U GGGCAAGCGG 1774
Arteriviridae Arterivirus Equine arteritis virus -1 NC_002532 CAGUGAAUCA GUUAAA C UGAGAGCGCC 5405
Porcine reproductive and respiratory syndrome virus -2 NC_001961 UGAGCCGUCA GGUUU UU GACCUCGUCU 3887-3888
Coronaviridae Alphacoronavirus Human coronavirus 229E -1 NC_002645 AUAACAGUUA UUUAAA C GAGUCCGGGG 12520
Betacoronavirus Human SARS coronavirus -1 NC_004718 CAUCAACGUU UUUAAA C GGGUUUGCGG 13398
Gammacoronavirus Avian Infectious bronchitis virus -1 NC_002645 AUAAGAAUUA UUUAAA C GGGUACGGGG 12354
Bafinivirus White bream virus 1 NC_008516 AGCAGCCGCC UUUAAA C TGGAGGGCAG 14555
Torovirus Breda virus 1 NC_007447 CUGUUGGUGA UUUAAA C UGCUGAGAGA 14184
FLaviviridae Flavivirus West nile virus -1 NC_009942 UAUGAUUGAC CCUUUU C AGUUGGGCCU 3552 30-50%
Mesoniviridae Mesonivirus Alphamesonivirus 1 -1 NC_015668 GCACCCUUUG GAUUUU C CAAAGUGGGC 7841
Picornaviridae Cardiovirus Encephalomyocardidis virus -1 NC_001479 CAAGGAAACA GGUUUU C CAGACCCAAG 3998
Tombusviridae Dianthovirus Red clover necrotic mosaic virus -1 NC_003756 AAUCCCUUGA GGAUUU U UAGGCGGCGG 831
Retroviridae Alpharetrovirus Avian leukosis virus -1 NC_001408 UCCGCUUGAC AAAUUU A UAGGGAGGGC 2474
Betaretrovirus Mouse mammary tumor virus -1 NC_001503 CUGAAAAUUC AAAAAA C UUGUAAAGGG 2082
Deltaretrovirus Human T-lymphotropic virus 1 -1 NC_001436 UCCCACACCC AAAAAA C UCCAUAGGGG 1718
Lentivirus Human immunodeficiency virus 1 -1 NC_001802 GACAGGCUAA UUUUUU A GGGAAGAUCU 1637
Unclassified Sobemovirus Cocksfoot mottle sobemovirus -1 NC_002618 CAAUCCGGCC UUUAAA C UACCAGCGGG 1640
Umbravirus Groundnut rosette virus -1 NC_003603 CCGGGGCACA AAAUUU U UAGUUGGGGA 867
Myoviridae Mulikevirus Mu phage -2 NC_000929 CGCUGACACA AGAAC GG GGGCGAGUGG 23960
Myoviridae P2likeviruses Enterobacteria phage P2 -1 NC_003603 CGGUAAGGUG GUCGGU U UUUUGUCGCC 19679
Siphoviridae Lambdalikevirus Phage lambda +1 NC_001416 AGAGCCUGUU UCUGC GG GAAAGTGTTC 10110
Togaviridae Alphavirus Sindbis virus +1 NC_001547 GCUGCCUGCC UUUUUU A GUGGUUCCGG 10028 10-18%

Source:Michael Bekaert, thesis (in french) etude du decalage de phase de lecture dans le genome de Saccharomyces cerevisiae

+1 ribosomal frameshift

This frameshift is mechanistically different from the -1. The ribosome stalls on a slippery sequence, making a long pause at a rare codon for which few tRNAs are available. This long pause can be resolved by a movement forward of one nucleotide, creating one or more mismatchs between tRNAs and mRNA. Translation resolves on the advanced ribosome in the +1 frame.

Family Genus Virus Type RefSeq .......... XXXYYY Z .......... Frameshift position Freq Ref
+1 FRAMESHIFTS (rare codon in yellow)
Orthomyxoviridae Influenzavirus A Influenzavirus A +1 NC_002022 UCUGGGAUUC CUUU CGU CAGUCCGAG 599
Totiviridae Leishmaniavirus Leishmania RNA virus 1-4 +1 NC_003601 Not Identified NA
Siphoviridae Lambdalikevirus Bacillus phage SPP1 +1 NC_004166 ACTGTTCCCG CTC CCC TAATGCGCC 11,111 ~10%

Matching UniProtKB/Swiss-Prot entries

544 entries grouped by protein (browse by keywords)

All proteins are shown (view only 405 complete proteome entries)

1 entry

Truncated polyprotein 1aTF

1ATF_PRRSS
Porcine reproductive and respiratory syndrome virus (isolate Pig/United States/SD 01-08/2001) (PRRSV)

3 entries

Protein 2B*

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ALT2B_EMCVR
Encephalomyocarditis virus (strain Rueckert) (EMCV) reference strain
ALT2B_EMCV
Encephalomyocarditis virus
ALT2B_ENMGO
Mengo encephalomyocarditis virus

1 entry

Major capsid protein 10A (Gene product 10A) (Gp10A) (Major head protein 10A)

CAPSA_BPT7
Enterobacteria phage T7 (Bacteriophage T7) reference strain

1 entry

Minor capsid protein 10B (Gene product 10B) (Gp10B) (Minor head protein 10B)

CAPSB_BPT7
Enterobacteria phage T7 (Bacteriophage T7) reference strain

2 entries

Capsid readthrough protein (CP-RT)

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CAPSD_BNYVS
Beet necrotic yellow vein virus (isolate Japan/S) (BNYVV) reference strain
CAPSD_GLVWB
Giardia lamblia virus (isolate Wang) (GLV) reference strain

Major capsid protein (Gag protein) (Major coat protein)

2 entries

F protein (Alternate reading frame protein/F-protein) (ARFP/F) (Frameshifted protein) (p16) (p17)

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F_HCVH
Hepatitis C virus genotype 1a (isolate H) (HCV) reference strain
F_HCV1
Hepatitis C virus genotype 1a (isolate 1) (HCV)

2 entries

Tail assembly protein G (Gene product 14) (gp14) (Minor tail protein G) (Tail assembly chaperone) (TAC)

Select_allDeselect_all
G_BPN15
Escherichia phage N15 (Bacteriophage N15) reference strain
G_LAMBD
Escherichia phage lambda (Bacteriophage lambda) reference strain

Tail assembly protein G (Gene product G) (gpG) (Minor tail protein G) (Tail assembly chaperone) (TAC)

79 entries

Gag polyprotein (Pr55Gag)

Select_allDeselect_all
GAG_HTL1C
Human T-cell leukemia virus 1 (isolate Caribbea HS-35 subtype A) (HTLV-1) reference strain

Gag polyprotein (Pr53Gag)

GAG_HTL32
Human T-cell leukemia virus 3 (strain 2026ND) (HTLV-3) reference strain

Gag polyprotein (Pr53Gag)

GAG_HTLV2
Human T-cell leukemia virus 2 (HTLV-2) reference strain

Gag polyprotein (Pr53Gag)

GAG_HV1B1
Human immunodeficiency virus type 1 group M subtype B (isolate BH10) (HIV-1) reference strain
GAG_HV1BR
Human immunodeficiency virus type 1 group M subtype B (isolate BRU/LAI) (HIV-1) reference strain
GAG_HV1EL
Human immunodeficiency virus type 1 group M subtype D (isolate ELI) (HIV-1) reference strain
GAG_HV1H2
Human immunodeficiency virus type 1 group M subtype B (isolate HXB2) (HIV-1) reference strain
GAG_HV1MA
Human immunodeficiency virus type 1 group M subtype A (isolate MAL) (HIV-1) reference strain
GAG_HV1MN
Human immunodeficiency virus type 1 group M subtype B (isolate MN) (HIV-1) reference strain
GAG_HV1VI
Human immunodeficiency virus type 1 group M subtype F1 (isolate VI850) (HIV-1) reference strain
GAG_HV1YF
Human immunodeficiency virus type 1 group N (isolate YBF30) (HIV-1) reference strain
GAG_HV2BE
Human immunodeficiency virus type 2 subtype A (isolate BEN) (HIV-2) reference strain
GAG_HV2ST
Human immunodeficiency virus type 2 subtype A (isolate ST) (HIV-2) reference strain
GAG_MMTVC
Mouse mammary tumor virus (strain C3H) (MMTV) reference strain

Gag polyprotein

GAG_RSVSA
Avian leukosis virus RSA (RSV-SRA) (Rous sarcoma virus (strain Schmidt-Ruppin A)) reference strain

Gag-Pro polyprotein

GAG_SCVLA
Saccharomyces cerevisiae virus L-A (ScV-L-A) (ScVL1) reference strain

Major capsid protein (Gag protein) (Major coat protein)

GAG_SIVEK
Simian immunodeficiency virus (isolate EK505) (SIV-cpz) (Chimpanzee immunodeficiency virus) reference strain
GAG_SIVMB
Simian immunodeficiency virus (isolate MB66) (SIV-cpz) (Chimpanzee immunodeficiency virus) reference strain
GAG_SIVTN
Simian immunodeficiency virus (isolate TAN1) (SIV-cpz) (Chimpanzee immunodeficiency virus) reference strain
GAG_BIV29
Bovine immunodeficiency virus (strain R29) (BIV) (Bovine immunodeficiency-like virus)

Gag polyprotein (Pr53Gag)

GAG_HTL1A
Human T-cell leukemia virus 1 (strain Japan ATK-1 subtype A) (HTLV-1)

Gag polyprotein (Pr53Gag)

GAG_HTL1L
Human T-cell leukemia virus 1 (isolate Melanesia mel5 subtype C) (HTLV-1)

Gag polyprotein (Pr53Gag)

GAG_HTL1M
Human T-cell leukemia virus 1 (strain Japan MT-2 subtype A) (HTLV-1)

Gag polyprotein (Pr53Gag)

GAG_HTL3P
Human T-cell leukemia virus 3 (strain Pyl43) (HTLV-3)

Gag polyprotein (Pr53Gag)

GAG_HV190
Human immunodeficiency virus type 1 group M subtype H (isolate 90CF056) (HIV-1)
GAG_HV192
Human immunodeficiency virus type 1 group M subtype C (isolate 92BR025) (HIV-1)
GAG_HV193
Human immunodeficiency virus type 1 group M subtype F1 (isolate 93BR020) (HIV-1)
GAG_HV196
Human immunodeficiency virus type 1 group M subtype K (isolate 96CM-MP535) (HIV-1)
GAG_HV197
Human immunodeficiency virus type 1 group M subtype K (isolate 97ZR-EQTB11) (HIV-1)
GAG_HV19N
Human immunodeficiency virus type 1 group M subtype G (isolate 92NG083) (HIV-1)
GAG_HV1A2
Human immunodeficiency virus type 1 group M subtype B (isolate ARV2/SF2) (HIV-1)
GAG_HV1AN
Human immunodeficiency virus type 1 group O (isolate ANT70) (HIV-1)
GAG_HV1B5
Human immunodeficiency virus type 1 group M subtype B (isolate BH5) (HIV-1)
GAG_HV1B9
Human immunodeficiency virus type 1 group M subtype B (strain 89.6) (HIV-1)
GAG_HV1C4
Human immunodeficiency virus type 1 group M subtype B (isolate CDC-451) (HIV-1)
GAG_HV1ET
Human immunodeficiency virus type 1 group M subtype C (isolate ETH2220) (HIV-1)
GAG_HV1J3
Human immunodeficiency virus type 1 group M subtype B (isolate JH32) (HIV-1)
GAG_HV1JR
Human immunodeficiency virus type 1 group M subtype B (isolate JRCSF) (HIV-1)
GAG_HV1LW
Human immunodeficiency virus type 1 group M subtype B (isolate LW123) (HIV-1)
GAG_HV1M2
Human immunodeficiency virus type 1 group M subtype F2 (isolate MP257) (HIV-1)
GAG_HV1MP
Human immunodeficiency virus type 1 group M subtype F2 (isolate MP255) (HIV-1)
GAG_HV1MV
Human immunodeficiency virus type 1 group O (isolate MVP5180) (HIV-1)
GAG_HV1N5
Human immunodeficiency virus type 1 group M subtype B (isolate NY5) (HIV-1)
GAG_HV1ND
Human immunodeficiency virus type 1 group M subtype D (isolate NDK) (HIV-1)
GAG_HV1OY
Human immunodeficiency virus type 1 group M subtype B (isolate OYI) (HIV-1)
GAG_HV1RH
Human immunodeficiency virus type 1 group M subtype B (isolate RF/HAT3) (HIV-1)
GAG_HV1S2
Human immunodeficiency virus type 1 group M subtype J (isolate SE9280) (HIV-1)
GAG_HV1S9
Human immunodeficiency virus type 1 group M subtype J (isolate SE9173) (HIV-1)
GAG_HV1SE
Human immunodeficiency virus type 1 group M subtype G (isolate SE6165) (HIV-1)
GAG_HV1U4
Human immunodeficiency virus type 1 group M subtype A (isolate U455) (HIV-1)
GAG_HV1V9
Human immunodeficiency virus type 1 group M subtype H (isolate VI991) (HIV-1)
GAG_HV1Y2
Human immunodeficiency virus type 1 group M subtype B (isolate YU-2) (HIV-1)
GAG_HV1YB
Human immunodeficiency virus type 1 group N (isolate YBF106) (HIV-1)
GAG_HV1Z2
Human immunodeficiency virus type 1 group M subtype D (isolate Z2/CDC-Z34) (HIV-1)
GAG_HV2CA
Human immunodeficiency virus type 2 subtype A (isolate CAM2) (HIV-2)
GAG_HV2D1
Human immunodeficiency virus type 2 subtype A (isolate D194) (HIV-2)
GAG_HV2D2
Human immunodeficiency virus type 2 subtype B (isolate D205) (HIV-2)
GAG_HV2EH
Human immunodeficiency virus type 2 subtype B (isolate EHO) (HIV-2)
GAG_HV2G1
Human immunodeficiency virus type 2 subtype A (isolate Ghana-1) (HIV-2)
GAG_HV2KR
Human immunodeficiency virus type 2 subtype A (isolate KR) (HIV-2)
GAG_HV2NZ
Human immunodeficiency virus type 2 subtype A (isolate NIH-Z) (HIV-2)
GAG_HV2RO
Human immunodeficiency virus type 2 subtype A (isolate ROD) (HIV-2)
GAG_HV2SB
Human immunodeficiency virus type 2 subtype A (isolate SBLISY) (HIV-2)
GAG_HV2UC
Human immunodeficiency virus type 2 subtype B (isolate UC1) (HIV-2)
GAG_JEMBR
Jembrana disease virus (JDV)

Gag polyprotein (Pr53Gag)

GAG_RSVP
Rous sarcoma virus (strain Prague C) (RSV-PrC)

Gag-Pro polyprotein

GAG_RSVSB
Rous sarcoma virus (strain Schmidt-Ruppin B) (RSV-SRB)

Gag-Pro polyprotein

GAG_SCVLB
Saccharomyces cerevisiae virus L-BC (ScV-L-BC) (ScVLa)

Major capsid protein (Gag protein) (Major coat protein)

GAG_SIVCZ
Simian immunodeficiency virus (isolate CPZ GAB1) (SIV-cpz) (Chimpanzee immunodeficiency virus)
GAG_SIVG1
Simian immunodeficiency virus agm.grivet (isolate AGM gr-1) (SIV-agm.gri) (Simian immunodeficiency virus African green monkey grivet)
GAG_SIVGB
Simian immunodeficiency virus (isolate GB1) (SIV-mnd) (Simian immunodeficiency virus mandrill)
GAG_SIVM1
Simian immunodeficiency virus (isolate Mm142-83) (SIV-mac) (Simian immunodeficiency virus rhesus monkey)
GAG_SIVMK
Simian immunodeficiency virus (isolate K6W) (SIV-mac) (Simian immunodeficiency virus rhesus monkey)
GAG_SIVMS
Simian immunodeficiency virus (isolate STM) (SIV-mac) (Simian immunodeficiency virus rhesus monkey)
GAG_SIVS4
Simian immunodeficiency virus (isolate F236/smH4) (SIV-sm) (Simian immunodeficiency virus sooty mangabey monkey)
GAG_SIVSP
Simian immunodeficiency virus (isolate PBj14/BCL-3) (SIV-sm) (Simian immunodeficiency virus sooty mangabey monkey)
GAG_SIVV1
Simian immunodeficiency virus agm.vervet (isolate AGM155) (SIV-agm.ver) (Simian immunodeficiency virus African green monkey vervet)
GAG_SIVVG
Simian immunodeficiency virus agm.vervet (isolate AGM3) (SIV-agm.ver) (Simian immunodeficiency virus African green monkey vervet)
GAG_SIVVT
Simian immunodeficiency virus agm.vervet (isolate AGM TYO-1) (SIV-agm.ver) (Simian immunodeficiency virus African green monkey vervet)

2 entries

Tail assembly protein GT (Gene product 15) (gp15) (Gene product GT) (gpGT) (Minor tail protein GT)

Select_allDeselect_all
GT_BPN15
Escherichia phage N15 (Bacteriophage N15) reference strain
GT_LAMBD
Escherichia phage lambda (Bacteriophage lambda) reference strain

Tail assembly protein GT (Gene product GT) (gpGT) (Minor tail protein GT)

24 entries

P3N-PIPO polyprotein

Select_allDeselect_all
MVP_PVYN
Potato virus Y (strain N) (PVY) reference strain
MVP_BCMNN
Bean common mosaic necrosis virus (strain NL-3) (BCMNV) (Bean common mosaic virus serotype A (strain NL-3))
MVP_BTMV
Beet mosaic virus (BtMV)
MVP_BYMV
Bean yellow mosaic virus
MVP_LMV0
Lettuce mosaic virus (strain 0 / isolate French) (LMV)
MVP_LMVE
Lettuce mosaic virus (strain E) (LMV)
MVP_PEMVC
Pepper mottle virus (isolate California) (PeMV) (PepMoV C)
MVP_PEMVM
Peanut mottle virus (strain M)
MVP_PPVD
Plum pox potyvirus (strain D) (PPV)
MVP_PPVNA
Plum pox potyvirus (isolate NAT) (PPV)
MVP_PPVSK
Plum pox potyvirus (strain SK 68) (PPV)
MVP_PRSVH
Papaya ringspot virus (strain P / mutant HA)
MVP_PSBMV
Pea seed-borne mosaic virus (strain DPD1)
MVP_PVMA
Potato virus A (PVA)
MVP_PVYHU
Potato virus Y (strain Hungarian) (PVY)
MVP_SBMVG
Soybean mosaic virus (strain G2) (SMV)
MVP_SBMVN
Soybean mosaic virus (strain N) (SMV)
MVP_TEV
Tobacco etch virus (TEV)
MVP_TUMVJ
Turnip mosaic virus (strain Japanese) (TuMV)
MVP_TUMVQ
Turnip mosaic virus (strain Quebec) (TuMV)
MVP_TVMV
Tobacco vein mottling virus (TVMV)
MVP_ZYMVC
Zucchini yellow mosaic virus (strain California) (ZYMV)
MVP_ZYMVR
Zucchini yellow mosaic virus (strain Reunion Island) (ZYMV)
MVP_ZYMVS
Zucchini yellow mosaic virus (strain Singapore) (ZYMV)

9 entries

Non-structural polyprotein 1A

Select_allDeselect_all
NS1A_HASV1
Human astrovirus-1 (HAstV-1) reference strain
NS1A_TASV1
Turkey astrovirus 1 (TAstV-1) reference strain
NS1A_ANV1
Avian nephritis virus 1 (ANV-1)
NS1A_HASV4
Human astrovirus-4 (HAstV-4)
NS1A_HASV5
Human astrovirus-5 (HAstV-5)
NS1A_HASV8
Human astrovirus-8 (HAstV-8)
NS1A_MASV1
Mink astrovirus 1 (MAstV-1)
NS1A_OASV1
Ovine astrovirus 1 (OAstV-1)
NS1A_TASV2
Turkey astrovirus 2 (TAstV-2)

9 entries

Non-structural polyprotein 1AB

Select_allDeselect_all
NS1AB_HASV1
Human astrovirus-1 (HAstV-1) reference strain
NS1AB_TASV1
Turkey astrovirus 1 (TAstV-1) reference strain
NS1AB_ANV1
Avian nephritis virus 1 (ANV-1)
NS1AB_HASV4
Human astrovirus-4 (HAstV-4)
NS1AB_HASV5
Human astrovirus-5 (HAstV-5)
NS1AB_HASV8
Human astrovirus-8 (HAstV-8)
NS1AB_MASV1
Mink astrovirus 1 (MAstV-1)
NS1AB_OASV1
Ovine astrovirus 1 (OAstV-1)
NS1AB_TASV2
Turkey astrovirus 2 (TAstV-2)

5 entries

Protein P1 (69.7 kDa protein) (Genome-linked protein precursor) (Protein ORF1)

Select_allDeselect_all
P1_PEMVW
Pea enation mosaic virus-1 (strain WSG) (PEMV-1) reference strain

Protein P1 (Genome-linked protein precursor) (ORF1 protein)

P1_PLRV1
Potato leafroll virus (strain Potato/Scotland/strain 1/1984) (PLrV) reference strain
P1_PNLV
Poinsettia latent virus (isolate Euphorbia pulcherrima/Germany/Siepen/2005) (PnLV) (Poinsettia cryptic virus) reference strain

Genome-linked protein precursor

P1_TYYVF
Turnip yellows virus (isolate FL-1) (TuYV) (BWYV-FL1) reference strain

Protein P1 (66.2 kDa protein) (Genome-linked protein precursor) (Protein ORF1)

P1_PLRVW
Potato leafroll virus (strain Potato/Netherlands/Wageningen/1989) (PLrV)

3 entries

Polyprotein P2A

Select_allDeselect_all
P2A_CFMVN
Cocksfoot mottle virus (isolate Dactylis glomerata/Norway/CfMV-NO/1995) (CfMV) reference strain
P2A_SBMVA
Southern bean mosaic virus (isolate Bean/United States/Arkansas) (SBMV) reference strain
P2A_SCPMV
Southern cowpea mosaic virus (SCPMV) (Southern bean mosaic virus (strain cowpea)) reference strain

96 entries

Polymerase acidic protein (EC 3.1.-.-) (RNA-directed RNA polymerase subunit P2)

Select_allDeselect_all
PA_I34A1
Influenza A virus (strain A/Puerto Rico/8/1934 H1N1) reference strain
PA_I57A5
Influenza A virus (strain A/Singapore/1/1957 H2N2) reference strain
PA_I68A4
Influenza A virus (strain A/Hong Kong/1/1968 H3N2) reference strain
PA_I77AF
Influenza A virus (strain A/Gull/Maryland/704/1977 H13N6) reference strain
PA_I96A0
Influenza A virus (strain A/Goose/Guangdong/1/1996 H5N1 genotype Gs/Gd) reference strain
PA_INBLE
Influenza B virus (strain B/Lee/1940) reference strain
PA_INCAA
Influenza C virus (strain C/Ann Arbor/1/1950) reference strain
PA_I000X
Influenza A virus (strain A/X-31 H3N2)
PA_I00A0
Influenza A virus (strain A/Duck/Hong Kong/2986.1/2000 H5N1 genotype C)
PA_I00A1
Influenza A virus (strain A/New Zealand:South Canterbury/35/2000 H1N1)
PA_I01A0
Influenza A virus (strain A/Silky Chicken/Hong Kong/SF189/2001 H5N1 genotype A)
PA_I01A3
Influenza A virus (strain A/Chicken/Hong Kong/715.5/2001 H5N1 genotype E)
PA_I02A0
Influenza A virus (strain A/Chicken/Brescia/1902 H7N7)
PA_I02A1
Influenza A virus (strain A/Guinea fowl/Hong Kong/38/2002 H5N1 genotype X0)
PA_I02A2
Influenza A virus (strain A/Chicken/Hong Kong/31.2/2002 H5N1 genotype X1)
PA_I02A4
Influenza A virus (strain A/Silky Chicken/Hong Kong/YU100/2002 H5N1 genotype X3)
PA_I02A5
Influenza A virus (strain A/Chicken/Hong Kong/96.1/2002 H5N1 genotype Y)
PA_I02A6
Influenza A virus (strain A/Chicken/Hong Kong/YU22/2002 H5N1 genotype Z)
PA_I05A1
Influenza A virus (strain A/Goose/Guangxi/345/2005 H5N1 genotype G)
PA_I06A0
Influenza A virus (strain A/Russia:St.Petersburg/8/2006 H1N1)
PA_I07A0
Influenza A virus (strain A/USA:Texas/UR06-0195/2007 H1N1)
PA_I18A0
Influenza A virus (strain A/Brevig Mission/1/1918 H1N1) (Influenza A virus (strain A/South Carolina/1/1918 H1N1))
PA_I30A0
Influenza A virus (strain A/Swine/Iowa/15/1930 H1N1)
PA_I33A0
Influenza A virus (strain A/Wilson-Smith/1933 H1N1) (Influenza A virus (strain A/WS/1933 H1N1))
PA_I34A0
Influenza A virus (strain A/Fowl plague virus/Rostock/8/1934 H7N1)
PA_I35A3
Influenza A virus (strain A/USA:Phila/1935 H1N1)
PA_I36A0
Influenza A virus (strain A/Henry/1936 H1N1)
PA_I40A0
Influenza A virus (strain A/Hickox/1940 H1N1)
PA_I43A0
Influenza A virus (strain A/USA:Iowa/1943 H1N1)
PA_I45A0
Influenza A virus (strain A/USA:Huston/AA/1945 H1N1)
PA_I47A0
Influenza A virus (strain A/Fort Monmouth/1/1947 H1N1)
PA_I49A1
Influenza A virus (strain A/Duck/Germany/1949 H10N7)
PA_I51A0
Influenza A virus (strain A/USA:Albany/12/1951 H1N1)
PA_I54A2
Influenza A virus (strain A/Malaysia:Malaya/302/1954 H1N1)
PA_I56A1
Influenza A virus (strain A/Duck/Czechoslovakia/1956 H4N6)
PA_I56A2
Influenza A virus (strain A/Duck/England/1/1956 H11N6)
PA_I57A0
Influenza A virus (strain A/Japan/305/1957 H2N2)
PA_I57A1
Influenza A virus (strain A/Leningrad/134/1957 H2N2)
PA_I57A2
Influenza A virus (strain A/Leningrad/134/17/1957 H2N2)
PA_I57A3
Influenza A virus (strain A/Leningrad/134/47/1957 H2N2)
PA_I59A0
Influenza A virus (strain A/Chicken/Scotland/1959 H5N1)
PA_I60A0
Influenza A virus (strain A/Ann Arbor/6/1960 H2N2)
PA_I61A1
Influenza A virus (strain A/Swine/Wisconsin/1/1961 H1N1)
PA_I66A1
Influenza A virus (strain A/Turkey/Wisconsin/1/1966 H9N2)
PA_I67A0
Influenza A virus (strain A/Tokyo/3/1967 H2N2)
PA_I67A2
Influenza A virus (strain A/Swine/Wisconsin/1/1967 H1N1)
PA_I68A3
Influenza A virus (strain A/Turkey/Ontario/6118/1968 H8N4)
PA_I68A5
Influenza A virus (strain A/Korea/426/1968 H2N2)
PA_I68A6
Influenza A virus (strain A/Northern Territory/60/1968 H3N2) (Influenza A virus (strain NT60)) (Influenza A virus (strain A/NT/60/1968 H3N2))
PA_I69A0
Influenza A virus (strain A/England/878/1969 H3N2)
PA_I70A0
Influenza A virus (strain A/Qu/7/1970 H3N2)
PA_I71A1
Influenza A virus (strain A/Memphis/1/1971 H3N2)
PA_I72A2
Influenza A virus (strain A/Udorn/307/1972 H3N2)
PA_I72A3
Influenza A virus (strain A/Memphis/102/1972 H3N2)
PA_I72A4
Influenza A virus (strain A/Memphis/101/1972 H3N2)
PA_I73A4
Influenza A virus (strain A/Equine/London/1416/1973 H7N7)
PA_I73A5
Influenza A virus (strain A/Port Chalmers/1/1973 H3N2)
PA_I75A0
Influenza A virus (strain A/Beijing/39/1975 H3N2)
PA_I75A3
Influenza A virus (strain A/Victoria/3/1975 H3N2)
PA_I76A6
Influenza A virus (strain A/Memphis/110/1976 H3N2)
PA_I77A4
Influenza A virus (strain A/Swine/Colorado/1/1977 H3N2)
PA_I77AA
Influenza A virus (strain A/Brazil/11/1978 H1N1)
PA_I77AB
Influenza A virus (strain A/USSR/90/1977 H1N1)
PA_I77AD
Influenza A virus (strain A/Swine/Tennessee/26/1977 H1N1)
PA_I78A3
Influenza A virus (strain A/Mallard/New York/6750/1978 H2N2)
PA_I78A7
Influenza A virus (strain A/Memphis/2/1978 H3N2)
PA_I78A8
Influenza A virus (strain A/Memphis/18/1978 H3N2)
PA_I78A9
Influenza A virus (strain A/Swine/Hong Kong/81/1978 H3N2)
PA_I78AC
Influenza A virus (strain A/Turkey/Minnesota/501/1978 H6N8)
PA_I79A1
Influenza A virus (strain A/Pintail/Alberta/119/1979 H4N6)
PA_I79A7
Influenza A virus (strain A/Grey teal/Australia/2/1979 H4N4)
PA_I80A2
Influenza A virus (strain A/Seal/Massachusetts/1/1980 H7N7)
PA_I80A4
Influenza A virus (strain A/Memphis/4/1980 H3N2)
PA_I80A6
Influenza A virus (strain A/Duck/Hokkaido/8/1980 H3N8)
PA_I80A8
Influenza A virus (strain A/Turkey/Minnesota/833/1980 H4N2)
PA_I80AA
Influenza A virus (strain A/India/6263/1980 H1N1)
PA_I80AD
Influenza A virus (strain A/Gull/Minnesota/945/1980 H13N6)
PA_I82A4
Influenza A virus (strain A/Swine/Hong Kong/126/1982 H3N2)
PA_I83A1
Influenza A virus (strain A/Chile/1/1983 H1N1)
PA_I83A4
Influenza A virus (strain A/Turkey/Ireland/1378/1983 H5N8)
PA_I83A5
Influenza A virus (strain A/Chicken/Pennsylvania/1/1983 H5N2)
PA_I83A8
Influenza A virus (strain A/Hong Kong/5/1983 H3N2)
PA_I85A3
Influenza A virus (strain A/Chicken/Victoria/1/1985 H7N7)
PA_I85A7
Influenza A virus (strain A/Ruddy Turnstone/New Jersey/47/1985 H4N6)
PA_I86A3
Influenza A virus (strain A/Equine/Tennessee/5/1986 H3N8)
PA_I93A0
Influenza A virus (strain A/Kitakyushu/159/1993 H3N2)
PA_I96A2
Influenza A virus (strain A/USA:Memphis/10/1996 H1N1)
PA_I96A3
Influenza A virus (strain A/China:Nanchang/11/1996 H1N1)
PA_I97A0
Influenza A virus (strain A/Chicken/Hong Kong/220/1997 H5N1 genotype Gs/Gd)
PA_I97A1
Influenza A virus (strain A/Hong Kong/156/1997 H5N1 genotype Gs/Gd)
PA_INBAC
Influenza B virus (strain B/Ann Arbor/1/1966 [cold-adapted])
PA_INBAD
Influenza B virus (strain B/Ann Arbor/1/1966 [wild-type])
PA_INBP9
Influenza B virus (strain B/Panama/45/1990)
PA_INBSI
Influenza B virus (strain B/Singapore/222/1979)
PA_INCJH
Influenza C virus (strain C/Johannesburg/1/1966)
PA_INCJJ
Influenza C virus (strain C/JJ/1950)

87 entries

Protein PA-X

Select_allDeselect_all
PAX_I34A1
Influenza A virus (strain A/Puerto Rico/8/1934 H1N1) reference strain
PAX_I68A4
Influenza A virus (strain A/Hong Kong/1/1968 H3N2) reference strain
PAX_I77AF
Influenza A virus (strain A/Gull/Maryland/704/1977 H13N6) reference strain
PAX_I96A0
Influenza A virus (strain A/Goose/Guangdong/1/1996 H5N1 genotype Gs/Gd) reference strain
PAX_I000X
Influenza A virus (strain A/X-31 H3N2)
PAX_I00A0
Influenza A virus (strain A/Duck/Hong Kong/2986.1/2000 H5N1 genotype C)
PAX_I00A1
Influenza A virus (strain A/New Zealand:South Canterbury/35/2000 H1N1)
PAX_I01A0
Influenza A virus (strain A/Silky Chicken/Hong Kong/SF189/2001 H5N1 genotype A)
PAX_I01A3
Influenza A virus (strain A/Chicken/Hong Kong/715.5/2001 H5N1 genotype E)
PAX_I02A0
Influenza A virus (strain A/Chicken/Brescia/1902 H7N7)
PAX_I02A1
Influenza A virus (strain A/Guinea fowl/Hong Kong/38/2002 H5N1 genotype X0)
PAX_I02A2
Influenza A virus (strain A/Chicken/Hong Kong/31.2/2002 H5N1 genotype X1)
PAX_I02A4
Influenza A virus (strain A/Silky Chicken/Hong Kong/YU100/2002 H5N1 genotype X3)
PAX_I02A5
Influenza A virus (strain A/Chicken/Hong Kong/96.1/2002 H5N1 genotype Y)
PAX_I02A6
Influenza A virus (strain A/Chicken/Hong Kong/YU22/2002 H5N1 genotype Z)
PAX_I05A1
Influenza A virus (strain A/Goose/Guangxi/345/2005 H5N1 genotype G)
PAX_I06A0
Influenza A virus (strain A/Russia:St.Petersburg/8/2006 H1N1)
PAX_I07A0
Influenza A virus (strain A/USA:Texas/UR06-0195/2007 H1N1)
PAX_I18A0
Influenza A virus (strain A/Brevig Mission/1/1918 H1N1) (Influenza A virus (strain A/South Carolina/1/1918 H1N1))
PAX_I30A0
Influenza A virus (strain A/Swine/Iowa/15/1930 H1N1)
PAX_I33A0
Influenza A virus (strain A/Wilson-Smith/1933 H1N1) (Influenza A virus (strain A/WS/1933 H1N1))
PAX_I34A0
Influenza A virus (strain A/Fowl plague virus/Rostock/8/1934 H7N1)
PAX_I35A3
Influenza A virus (strain A/USA:Phila/1935 H1N1)
PAX_I36A0
Influenza A virus (strain A/Henry/1936 H1N1)
PAX_I40A0
Influenza A virus (strain A/Hickox/1940 H1N1)
PAX_I43A0
Influenza A virus (strain A/USA:Iowa/1943 H1N1)
PAX_I45A0
Influenza A virus (strain A/USA:Huston/AA/1945 H1N1)
PAX_I47A0
Influenza A virus (strain A/Fort Monmouth/1/1947 H1N1)
PAX_I49A1
Influenza A virus (strain A/Duck/Germany/1949 H10N7)
PAX_I51A0
Influenza A virus (strain A/USA:Albany/12/1951 H1N1)
PAX_I54A2
Influenza A virus (strain A/Malaysia:Malaya/302/1954 H1N1)
PAX_I56A1
Influenza A virus (strain A/Duck/Czechoslovakia/1956 H4N6)
PAX_I56A2
Influenza A virus (strain A/Duck/England/1/1956 H11N6)
PAX_I57A0
Influenza A virus (strain A/Japan/305/1957 H2N2)
PAX_I57A1
Influenza A virus (strain A/Leningrad/134/1957 H2N2)
PAX_I57A2
Influenza A virus (strain A/Leningrad/134/17/1957 H2N2)
PAX_I57A3
Influenza A virus (strain A/Leningrad/134/47/1957 H2N2)
PAX_I57A5
Influenza A virus (strain A/Singapore/1/1957 H2N2)
PAX_I59A0
Influenza A virus (strain A/Chicken/Scotland/1959 H5N1)
PAX_I60A0
Influenza A virus (strain A/Ann Arbor/6/1960 H2N2)
PAX_I61A1
Influenza A virus (strain A/Swine/Wisconsin/1/1961 H1N1)
PAX_I66A1
Influenza A virus (strain A/Turkey/Wisconsin/1/1966 H9N2)
PAX_I67A0
Influenza A virus (strain A/Tokyo/3/1967 H2N2)
PAX_I67A2
Influenza A virus (strain A/Swine/Wisconsin/1/1967 H1N1)
PAX_I68A3
Influenza A virus (strain A/Turkey/Ontario/6118/1968 H8N4)
PAX_I68A5
Influenza A virus (strain A/Korea/426/1968 H2N2)
PAX_I68A6
Influenza A virus (strain A/Northern Territory/60/1968 H3N2) (Influenza A virus (strain NT60)) (Influenza A virus (strain A/NT/60/1968 H3N2))
PAX_I69A0
Influenza A virus (strain A/England/878/1969 H3N2)
PAX_I70A0
Influenza A virus (strain A/Qu/7/1970 H3N2)
PAX_I71A1
Influenza A virus (strain A/Memphis/1/1971 H3N2)
PAX_I72A2
Influenza A virus (strain A/Udorn/307/1972 H3N2)
PAX_I72A3
Influenza A virus (strain A/Memphis/102/1972 H3N2)
PAX_I72A4
Influenza A virus (strain A/Memphis/101/1972 H3N2)
PAX_I73A4
Influenza A virus (strain A/Equine/London/1416/1973 H7N7)
PAX_I73A5
Influenza A virus (strain A/Port Chalmers/1/1973 H3N2)
PAX_I75A0
Influenza A virus (strain A/Beijing/39/1975 H3N2)
PAX_I76A6
Influenza A virus (strain A/Memphis/110/1976 H3N2)
PAX_I77A4
Influenza A virus (strain A/Swine/Colorado/1/1977 H3N2)
PAX_I77AA
Influenza A virus (strain A/Brazil/11/1978 H1N1)
PAX_I77AB
Influenza A virus (strain A/USSR/90/1977 H1N1)
PAX_I77AD
Influenza A virus (strain A/Swine/Tennessee/26/1977 H1N1)
PAX_I78A3
Influenza A virus (strain A/Mallard/New York/6750/1978 H2N2)
PAX_I78A7
Influenza A virus (strain A/Memphis/2/1978 H3N2)
PAX_I78A8
Influenza A virus (strain A/Memphis/18/1978 H3N2)
PAX_I78A9
Influenza A virus (strain A/Swine/Hong Kong/81/1978 H3N2)
PAX_I78AC
Influenza A virus (strain A/Turkey/Minnesota/501/1978 H6N8)
PAX_I79A1
Influenza A virus (strain A/Pintail/Alberta/119/1979 H4N6)
PAX_I79A7
Influenza A virus (strain A/Grey teal/Australia/2/1979 H4N4)
PAX_I80A2
Influenza A virus (strain A/Seal/Massachusetts/1/1980 H7N7)
PAX_I80A4
Influenza A virus (strain A/Memphis/4/1980 H3N2)
PAX_I80A6
Influenza A virus (strain A/Duck/Hokkaido/8/1980 H3N8)
PAX_I80A8
Influenza A virus (strain A/Turkey/Minnesota/833/1980 H4N2)
PAX_I80AA
Influenza A virus (strain A/India/6263/1980 H1N1)
PAX_I80AD
Influenza A virus (strain A/Gull/Minnesota/945/1980 H13N6)
PAX_I82A4
Influenza A virus (strain A/Swine/Hong Kong/126/1982 H3N2)
PAX_I83A1
Influenza A virus (strain A/Chile/1/1983 H1N1)
PAX_I83A4
Influenza A virus (strain A/Turkey/Ireland/1378/1983 H5N8)
PAX_I83A5
Influenza A virus (strain A/Chicken/Pennsylvania/1/1983 H5N2)
PAX_I83A8
Influenza A virus (strain A/Hong Kong/5/1983 H3N2)
PAX_I85A3
Influenza A virus (strain A/Chicken/Victoria/1/1985 H7N7)
PAX_I85A7
Influenza A virus (strain A/Ruddy Turnstone/New Jersey/47/1985 H4N6)
PAX_I86A3
Influenza A virus (strain A/Equine/Tennessee/5/1986 H3N8)
PAX_I93A0
Influenza A virus (strain A/Kitakyushu/159/1993 H3N2)
PAX_I96A2
Influenza A virus (strain A/USA:Memphis/10/1996 H1N1)
PAX_I96A3
Influenza A virus (strain A/China:Nanchang/11/1996 H1N1)
PAX_I97A0
Influenza A virus (strain A/Chicken/Hong Kong/220/1997 H5N1 genotype Gs/Gd)
PAX_I97A1
Influenza A virus (strain A/Hong Kong/156/1997 H5N1 genotype Gs/Gd)

73 entries

Gag-Pol polyprotein (Pr160Gag-Pol)

Select_allDeselect_all
POL_HTL1C
Human T-cell leukemia virus 1 (isolate Caribbea HS-35 subtype A) (HTLV-1) reference strain

Gag-Pro-Pol polyprotein (Pr160Gag-Pro-Pol)

POL_HTL32
Human T-cell leukemia virus 3 (strain 2026ND) (HTLV-3) reference strain

Gag-Pro-Pol polyprotein (Pr160Gag-Pro-Pol)

POL_HTLV2
Human T-cell leukemia virus 2 (HTLV-2) reference strain

Gag-Pro-Pol polyprotein (Pr160Gag-Pro-Pol)

POL_HV1B1
Human immunodeficiency virus type 1 group M subtype B (isolate BH10) (HIV-1) reference strain

pdb 1A9M; 1AJV; 1AJX; 1AXA; 1BQM; 1BQN; 1D4H; 1D4I; 1D4J; 1DLO; 1DW6; 1EBK; 1EBW; 1EBY; 1EBZ; 1EC0; 1EC1; 1EC2; 1EC3; 1EET; 1G35; 1GNM; 1GNN; 1GNO; 1HAR; 1HBV; 1HEF; 1HEG; 1HIH; 1HMV; 1HNI; 1HNV; 1HOS; 1HPS; 1HPZ; 1HQE; 1HQU; 1HRH; 1HTE; 1HTF; 1HTG; 1HVI; 1HVK; 1HVP; 1HVU; 1HYS; 1IKV; 1IKW; 1IKX; 1IKY; 1J5O; 1KJH; 1MER; 1MES; 1MET; 1MEU; 1N5Y; 1N6Q; 1NPA; 1NPV; 1NPW; 1QE1; 1QMC; 1R0A; 1RDH; 1RTD; 1RVL; 1RVM; 1RVN; 1RVO; 1RVP; 1RVQ; 1RVR; 1S6P; 1S6Q; 1S9E; 1S9G; 1SBG; 1SUQ; 1SV5; 1T03; 1T05; 1T7K; 1TV6; 1TVR; 1UWB; 1W5V; 1W5W; 1W5X; 1W5Y; 1YT9; 1ZP8; 1ZPA; 1ZSF; 1ZSR; 2AQU; 2B5J; 2B6A; 2BAN; 2BBB; 2BE2; 2EXF; 2G69; 2HB3; 2HMI; 2HNZ; 2HS1; 2HS2; 2I4D; 2I4U; 2I4V; 2I4W; 2I4X; 2I5J; 2IAJ; 2IC3; 2IDW; 2IEO; 2JZW; 2L45; 2L46; 2L4L; 2UXZ; 2UY0; 2VG5; 2VG6; 2VG7; 2X4U; 2YKM; 2YKN; 2ZD1; 2ZE2; 3AVI; 3BGR; 3DLK; 3GGA; 3GGV; 3GGX; 3HVT; 3IG1; 3IRX; 3IS9; 3ISN; 3ITH; 3JSM; 3JYT; 3K2P; 3K4V; 3KLE; 3KLF; 3KLG; 3KLH; 3KLI; 3NDT; 3NU3; 3NU4; 3NU5; 3NU6; 3NU9; 3NUJ; 3NUO; 3OK9; 3PSU; 3QAA; 3QLH; 3QO9; 3TKG; 3TKW; 3TL9; 3TLH; 3V4I; 3V6D; 3V81; 3ZPS; 3ZPT; 3ZPU; 4COE; 4CP7; 4CPQ; 4CPR; 4CPS; 4CPT; 4CPU; 4CPW; 4CPX; 4DG1; 4G1Q; 4G8G; 4G8I; 4G9D; 4G9F; 4H4M; 4H4O; 4I2P; 4I2Q; 4ICL; 4ID5; 4IDK; 4IFV; 4IFY; 4IG0; 4IG3; 4KFB; 4KKO; 4KO0; 4LSL; 4LSN; 4MFB; 4O44; 4O4G; 4OJR; 4PQU; 4PUO; 4PWD; 4Q0B; 4QAG; 4R5P; 4RW4; 4RW6; 4RW7; 4RW8; 4RW9; 4U8W; 4WE1; 4YE3; 4YHQ; 4ZIP; 4ZLS; 5AGZ; 5AH6; 5AH7; 5AH8; 5AH9; 5AHA; 5AHB; 5AHC; 5BRY; 5BS4; 5C24; 5C25; 5C42; 5CYM; 5CYQ; 5D3G; 5FDL; 5HBM; 5HLF; 5HP1; 5HRO; 5I3U; 5I42; 5J1E; 5JFP; 5JFU; 5JG1; 5T6Z; 5T70; 5TEP; 5TER; 5TW3; 5TXL; 5TXM; 5TXN; 5TXO; 5TXP

POL_HV1BR
Human immunodeficiency virus type 1 group M subtype B (isolate BRU/LAI) (HIV-1) reference strain
POL_HV1EL
Human immunodeficiency virus type 1 group M subtype D (isolate ELI) (HIV-1) reference strain
POL_HV1H2
Human immunodeficiency virus type 1 group M subtype B (isolate HXB2) (HIV-1) reference strain
POL_HV1MA
Human immunodeficiency virus type 1 group M subtype A (isolate MAL) (HIV-1) reference strain
POL_HV1MN
Human immunodeficiency virus type 1 group M subtype B (isolate MN) (HIV-1) reference strain
POL_HV1VI
Human immunodeficiency virus type 1 group M subtype F1 (isolate VI850) (HIV-1) reference strain
POL_HV1YF
Human immunodeficiency virus type 1 group N (isolate YBF30) (HIV-1) reference strain
POL_HV2BE
Human immunodeficiency virus type 2 subtype A (isolate BEN) (HIV-2) reference strain
POL_HV2ST
Human immunodeficiency virus type 2 subtype A (isolate ST) (HIV-2) reference strain
POL_MMTVC
Mouse mammary tumor virus (strain C3H) (MMTV) reference strain

Gag-Pro-Pol polyprotein

POL_RSVSA
Avian leukosis virus RSA (RSV-SRA) (Rous sarcoma virus (strain Schmidt-Ruppin A)) reference strain

Gag-Pro-Pol polyprotein

POL_SIVEK
Simian immunodeficiency virus (isolate EK505) (SIV-cpz) (Chimpanzee immunodeficiency virus) reference strain
POL_SIVMB
Simian immunodeficiency virus (isolate MB66) (SIV-cpz) (Chimpanzee immunodeficiency virus) reference strain
POL_SIVTN
Simian immunodeficiency virus (isolate TAN1) (SIV-cpz) (Chimpanzee immunodeficiency virus) reference strain
POL_BIV29
Bovine immunodeficiency virus (strain R29) (BIV) (Bovine immunodeficiency-like virus)

Gag-Pol polyprotein (Pr170Gag-Pol)

POL_HTL1A
Human T-cell leukemia virus 1 (strain Japan ATK-1 subtype A) (HTLV-1)

Gag-Pro-Pol polyprotein (Pr160Gag-Pro-Pol)

POL_HTL1L
Human T-cell leukemia virus 1 (isolate Melanesia mel5 subtype C) (HTLV-1)

Gag-Pro-Pol polyprotein (Pr160Gag-Pro-Pol)

POL_HTL3P
Human T-cell leukemia virus 3 (strain Pyl43) (HTLV-3)

Gag-Pro-Pol polyprotein (Pr160Gag-Pro-Pol)

POL_HV190
Human immunodeficiency virus type 1 group M subtype H (isolate 90CF056) (HIV-1)
POL_HV192
Human immunodeficiency virus type 1 group M subtype C (isolate 92BR025) (HIV-1)
POL_HV193
Human immunodeficiency virus type 1 group M subtype F1 (isolate 93BR020) (HIV-1)
POL_HV196
Human immunodeficiency virus type 1 group M subtype K (isolate 96CM-MP535) (HIV-1)
POL_HV197
Human immunodeficiency virus type 1 group M subtype K (isolate 97ZR-EQTB11) (HIV-1)
POL_HV19N
Human immunodeficiency virus type 1 group M subtype G (isolate 92NG083) (HIV-1)
POL_HV1A2
Human immunodeficiency virus type 1 group M subtype B (isolate ARV2/SF2) (HIV-1)
POL_HV1AN
Human immunodeficiency virus type 1 group O (isolate ANT70) (HIV-1)
POL_HV1B5
Human immunodeficiency virus type 1 group M subtype B (isolate BH5) (HIV-1)
POL_HV1B9
Human immunodeficiency virus type 1 group M subtype B (strain 89.6) (HIV-1)
POL_HV1ET
Human immunodeficiency virus type 1 group M subtype C (isolate ETH2220) (HIV-1)
POL_HV1JR
Human immunodeficiency virus type 1 group M subtype B (isolate JRCSF) (HIV-1)
POL_HV1LW
Human immunodeficiency virus type 1 group M subtype B (isolate LW123) (HIV-1)
POL_HV1M2
Human immunodeficiency virus type 1 group M subtype F2 (isolate MP257) (HIV-1)
POL_HV1MP
Human immunodeficiency virus type 1 group M subtype F2 (isolate MP255) (HIV-1)
POL_HV1MV
Human immunodeficiency virus type 1 group O (isolate MVP5180) (HIV-1)
POL_HV1N5
Human immunodeficiency virus type 1 group M subtype B (isolate NY5) (HIV-1)
POL_HV1ND
Human immunodeficiency virus type 1 group M subtype D (isolate NDK) (HIV-1)
POL_HV1OY
Human immunodeficiency virus type 1 group M subtype B (isolate OYI) (HIV-1)
POL_HV1RH
Human immunodeficiency virus type 1 group M subtype B (isolate RF/HAT3) (HIV-1)
POL_HV1S2
Human immunodeficiency virus type 1 group M subtype J (isolate SE9280) (HIV-1)
POL_HV1S9
Human immunodeficiency virus type 1 group M subtype J (isolate SE9173) (HIV-1)
POL_HV1SE
Human immunodeficiency virus type 1 group M subtype G (isolate SE6165) (HIV-1)
POL_HV1U4
Human immunodeficiency virus type 1 group M subtype A (isolate U455) (HIV-1)
POL_HV1V9
Human immunodeficiency virus type 1 group M subtype H (isolate VI991) (HIV-1)
POL_HV1Y2
Human immunodeficiency virus type 1 group M subtype B (isolate YU-2) (HIV-1)
POL_HV1YB
Human immunodeficiency virus type 1 group N (isolate YBF106) (HIV-1)
POL_HV1Z2
Human immunodeficiency virus type 1 group M subtype D (isolate Z2/CDC-Z34) (HIV-1)
POL_HV2CA
Human immunodeficiency virus type 2 subtype A (isolate CAM2) (HIV-2)
POL_HV2D1
Human immunodeficiency virus type 2 subtype A (isolate D194) (HIV-2)
POL_HV2D2
Human immunodeficiency virus type 2 subtype B (isolate D205) (HIV-2)
POL_HV2EH
Human immunodeficiency virus type 2 subtype B (isolate EHO) (HIV-2)
POL_HV2G1
Human immunodeficiency virus type 2 subtype A (isolate Ghana-1) (HIV-2)
POL_HV2KR
Human immunodeficiency virus type 2 subtype A (isolate KR) (HIV-2)
POL_HV2NZ
Human immunodeficiency virus type 2 subtype A (isolate NIH-Z) (HIV-2)
POL_HV2RO
Human immunodeficiency virus type 2 subtype A (isolate ROD) (HIV-2)
POL_HV2SB
Human immunodeficiency virus type 2 subtype A (isolate SBLISY) (HIV-2)
POL_HV2UC
Human immunodeficiency virus type 2 subtype B (isolate UC1) (HIV-2)
POL_JEMBR
Jembrana disease virus (JDV)

Gag-Pol polyprotein (Pr170Gag-Pol)

POL_RSVP
Rous sarcoma virus (strain Prague C) (RSV-PrC)

Gag-Pro-Pol polyprotein

POL_RSVSB
Rous sarcoma virus (strain Schmidt-Ruppin B) (RSV-SRB)

Gag-Pro-Pol polyprotein

POL_SIVCZ
Simian immunodeficiency virus (isolate CPZ GAB1) (SIV-cpz) (Chimpanzee immunodeficiency virus)
POL_SIVG1
Simian immunodeficiency virus agm.grivet (isolate AGM gr-1) (SIV-agm.gri) (Simian immunodeficiency virus African green monkey grivet)
POL_SIVGB
Simian immunodeficiency virus (isolate GB1) (SIV-mnd) (Simian immunodeficiency virus mandrill)
POL_SIVM1
Simian immunodeficiency virus (isolate Mm142-83) (SIV-mac) (Simian immunodeficiency virus rhesus monkey)
POL_SIVMK
Simian immunodeficiency virus (isolate K6W) (SIV-mac) (Simian immunodeficiency virus rhesus monkey)
POL_SIVS4
Simian immunodeficiency virus (isolate F236/smH4) (SIV-sm) (Simian immunodeficiency virus sooty mangabey monkey)
POL_SIVSP
Simian immunodeficiency virus (isolate PBj14/BCL-3) (SIV-sm) (Simian immunodeficiency virus sooty mangabey monkey)
POL_SIVV1
Simian immunodeficiency virus agm.vervet (isolate AGM155) (SIV-agm.ver) (Simian immunodeficiency virus African green monkey vervet)
POL_SIVVG
Simian immunodeficiency virus agm.vervet (isolate AGM3) (SIV-agm.ver) (Simian immunodeficiency virus African green monkey vervet)
POL_SIVVT
Simian immunodeficiency virus agm.vervet (isolate AGM TYO-1) (SIV-agm.ver) (Simian immunodeficiency virus African green monkey vervet)

29 entries

Genome polyprotein

Select_allDeselect_all
POLG_EMCVR
Encephalomyocarditis virus (strain Rueckert) (EMCV) reference strain
POLG_HCVH
Hepatitis C virus genotype 1a (isolate H) (HCV) reference strain
POLG_PVYN
Potato virus Y (strain N) (PVY) reference strain
POLG_BCMNN
Bean common mosaic necrosis virus (strain NL-3) (BCMNV) (Bean common mosaic virus serotype A (strain NL-3))
POLG_BTMV
Beet mosaic virus (BtMV)
POLG_BYMV
Bean yellow mosaic virus
POLG_EMCV
Encephalomyocarditis virus
POLG_ENMGO
Mengo encephalomyocarditis virus
POLG_HCV1
Hepatitis C virus genotype 1a (isolate 1) (HCV)
POLG_LMV0
Lettuce mosaic virus (strain 0 / isolate French) (LMV)
POLG_LMVE
Lettuce mosaic virus (strain E) (LMV)
POLG_PEMVC
Pepper mottle virus (isolate California) (PeMV) (PepMoV C)
POLG_PEMVM
Peanut mottle virus (strain M)
POLG_PPVD
Plum pox potyvirus (strain D) (PPV)
POLG_PPVNA
Plum pox potyvirus (isolate NAT) (PPV)
POLG_PPVSK
Plum pox potyvirus (strain SK 68) (PPV)
POLG_PRSVH
Papaya ringspot virus (strain P / mutant HA)
POLG_PSBMV
Pea seed-borne mosaic virus (strain DPD1)
POLG_PVMA
Potato virus A (PVA)
POLG_PVYHU
Potato virus Y (strain Hungarian) (PVY)
POLG_SBMVG
Soybean mosaic virus (strain G2) (SMV)
POLG_SBMVN
Soybean mosaic virus (strain N) (SMV)
POLG_TEV
Tobacco etch virus (TEV)
POLG_TUMVJ
Turnip mosaic virus (strain Japanese) (TuMV)
POLG_TUMVQ
Turnip mosaic virus (strain Quebec) (TuMV)
POLG_TVMV
Tobacco vein mottling virus (TVMV)
POLG_ZYMVC
Zucchini yellow mosaic virus (strain California) (ZYMV)
POLG_ZYMVR
Zucchini yellow mosaic virus (strain Reunion Island) (ZYMV)
POLG_ZYMVS
Zucchini yellow mosaic virus (strain Singapore) (ZYMV)

1 entry

Structural polyprotein (p130)

POLS_SFV
Semliki forest virus (SFV) reference strain

1 entry

Frameshifted structural polyprotein (p130)

POLSF_SFV
Semliki forest virus (SFV)

7 entries

Gag-Pro polyprotein (Pr76Gag-Pro)

Select_allDeselect_all
PRO_HTL1C
Human T-cell leukemia virus 1 (isolate Caribbea HS-35 subtype A) (HTLV-1) reference strain
PRO_HTL32
Human T-cell leukemia virus 3 (strain 2026ND) (HTLV-3) reference strain
PRO_HTLV2
Human T-cell leukemia virus 2 (HTLV-2) reference strain
PRO_MMTVC
Mouse mammary tumor virus (strain C3H) (MMTV) reference strain

Gag-Pro polyprotein

PRO_HTL1A
Human T-cell leukemia virus 1 (strain Japan ATK-1 subtype A) (HTLV-1)
PRO_HTL1L
Human T-cell leukemia virus 1 (isolate Melanesia mel5 subtype C) (HTLV-1)
PRO_HTL3P
Human T-cell leukemia virus 3 (strain Pyl43) (HTLV-3)

33 entries

Replicase polyprotein 1a (pp1a) (ORF1a polyprotein)

Select_allDeselect_all
R1A_BC512
Bat coronavirus 512/2005 (BtCoV) (BtCoV/512/2005) reference strain
R1A_BCHK4
Bat coronavirus HKU4 (BtCoV) (BtCoV/HKU4/2004) reference strain
R1A_BCHK5
Bat coronavirus HKU5 (BtCoV) (BtCoV/HKU5/2004) reference strain
R1A_BCHK9
Bat coronavirus HKU9 (BtCoV) (BtCoV/HKU9) reference strain
R1A_BEV
Berne virus (BEV) reference strain
R1A_BRV1
Breda virus 1 (BRV-1) reference strain
R1A_CVEMC
Human coronavirus EMC (isolate United Kingdom/H123990006/2012) (HCoV-EMC) reference strain
R1A_CVH22
Human coronavirus 229E (HCoV-229E) reference strain
R1A_CVHN1
Human coronavirus HKU1 (isolate N1) (HCoV-HKU1) reference strain
R1A_CVHNL
Human coronavirus NL63 (HCoV-NL63) reference strain
R1A_CVHOC
Human coronavirus OC43 (HCoV-OC43) reference strain
R1A_CVHSA
Human SARS coronavirus (SARS-CoV) (Severe acute respiratory syndrome coronavirus) reference strain
R1A_CVM2
Murine coronavirus (strain 2) (MHV-2) (Murine hepatitis virus) reference strain
R1A_CVMA5
Murine coronavirus (strain A59) (MHV-A59) (Murine hepatitis virus) reference strain
R1A_CVPPU
Porcine transmissible gastroenteritis coronavirus (strain Purdue) (TGEV) reference strain
R1A_GLRV3
Grapevine leafroll-associated virus 3 (isolate United States/NY1) (GLRaV-3) (Grapevine leafroll-associated closterovirus (isolate 109)) reference strain

Replicase protein 1a

R1A_IBVB
Avian infectious bronchitis virus (strain Beaudette) (IBV) reference strain
R1A_LIYV9
Lettuce infectious yellows virus (isolate United States/92) (LIYV) reference strain

Replicase polyprotein 1a

R1A_PEDV7
Porcine epidemic diarrhea virus (strain CV777) (PEDV) reference strain
R1A_WBV24
White bream virus (isolate Blicca bjoerkna L./Germany/DF24/00) (WBV) reference strain
R1A_BC133
Bat coronavirus 133/2005 (BtCoV) (BtCoV/133/2005)
R1A_BC279
Bat coronavirus 279/2005 (BtCoV) (BtCoV/279/2005)
R1A_BCHK3
Bat coronavirus HKU3 (BtCoV) (SARS-like coronavirus HKU3)
R1A_BCRP3
Bat coronavirus Rp3/2004 (BtCoV/Rp3/2004) (SARS-like coronavirus Rp3)
R1A_CVBEN
Bovine coronavirus (strain 98TXSF-110-ENT) (BCoV-ENT) (BCV)
R1A_CVBLU
Bovine coronavirus (strain 98TXSF-110-LUN) (BCoV-LUN) (BCV)
R1A_CVBM
Bovine coronavirus (strain Mebus) (BCoV) (BCV)
R1A_CVBQ
Bovine coronavirus (strain Quebec) (BCoV) (BCV)
R1A_CVHN2
Human coronavirus HKU1 (isolate N2) (HCoV-HKU1)
R1A_CVHN5
Human coronavirus HKU1 (isolate N5) (HCoV-HKU1)
R1A_CVMJH
Murine coronavirus (strain JHM) (MHV-JHM) (Murine hepatitis virus)
R1A_IBVBC
Avian infectious bronchitis virus (strain Beaudette CK) (IBV)
R1A_IBVM
Avian infectious bronchitis virus (strain M41) (IBV)

35 entries

Replicase polyprotein 1ab (pp1ab) (ORF1ab polyprotein)

Select_allDeselect_all
R1AB_BC512
Bat coronavirus 512/2005 (BtCoV) (BtCoV/512/2005) reference strain
R1AB_BCHK4
Bat coronavirus HKU4 (BtCoV) (BtCoV/HKU4/2004) reference strain
R1AB_BCHK5
Bat coronavirus HKU5 (BtCoV) (BtCoV/HKU5/2004) reference strain
R1AB_BCHK9
Bat coronavirus HKU9 (BtCoV) (BtCoV/HKU9) reference strain
R1AB_BEV
Berne virus (BEV) reference strain
R1AB_BRV1
Breda virus 1 (BRV-1) reference strain
R1AB_BYVU
Beet yellows virus (isolate Ukraine) (BYV) (Sugar beet yellows virus) reference strain

Replicase polyprotein 1ab

R1AB_CVEMC
Human coronavirus EMC (isolate United Kingdom/H123990006/2012) (HCoV-EMC) reference strain
R1AB_CVH22
Human coronavirus 229E (HCoV-229E) reference strain
R1AB_CVHN1
Human coronavirus HKU1 (isolate N1) (HCoV-HKU1) reference strain
R1AB_CVHNL
Human coronavirus NL63 (HCoV-NL63) reference strain
R1AB_CVHOC
Human coronavirus OC43 (HCoV-OC43) reference strain
R1AB_CVHSA
Human SARS coronavirus (SARS-CoV) (Severe acute respiratory syndrome coronavirus) reference strain
R1AB_CVM2
Murine coronavirus (strain 2) (MHV-2) (Murine hepatitis virus) reference strain
R1AB_CVMA5
Murine coronavirus (strain A59) (MHV-A59) (Murine hepatitis virus) reference strain
R1AB_CVPPU
Porcine transmissible gastroenteritis coronavirus (strain Purdue) (TGEV) reference strain
R1AB_FIPV
Feline coronavirus (strain FIPV WSU-79/1146) (FCoV) reference strain
R1AB_GLRV3
Grapevine leafroll-associated virus 3 (isolate United States/NY1) (GLRaV-3) (Grapevine leafroll-associated closterovirus (isolate 109)) reference strain

Replicase polyprotein 1ab

R1AB_IBVB
Avian infectious bronchitis virus (strain Beaudette) (IBV) reference strain
R1AB_LIYV9
Lettuce infectious yellows virus (isolate United States/92) (LIYV) reference strain

Replicase polyprotein 1ab

R1AB_PEDV7
Porcine epidemic diarrhea virus (strain CV777) (PEDV) reference strain
R1AB_WBV24
White bream virus (isolate Blicca bjoerkna L./Germany/DF24/00) (WBV) reference strain
R1AB_BC133
Bat coronavirus 133/2005 (BtCoV) (BtCoV/133/2005)
R1AB_BC279
Bat coronavirus 279/2005 (BtCoV) (BtCoV/279/2005)
R1AB_BCHK3
Bat coronavirus HKU3 (BtCoV) (SARS-like coronavirus HKU3)
R1AB_BCRP3
Bat coronavirus Rp3/2004 (BtCoV/Rp3/2004) (SARS-like coronavirus Rp3)
R1AB_CVBEN
Bovine coronavirus (strain 98TXSF-110-ENT) (BCoV-ENT) (BCV)
R1AB_CVBLU
Bovine coronavirus (strain 98TXSF-110-LUN) (BCoV-LUN) (BCV)
R1AB_CVBM
Bovine coronavirus (strain Mebus) (BCoV) (BCV)
R1AB_CVBQ
Bovine coronavirus (strain Quebec) (BCoV) (BCV)
R1AB_CVHN2
Human coronavirus HKU1 (isolate N2) (HCoV-HKU1)
R1AB_CVHN5
Human coronavirus HKU1 (isolate N5) (HCoV-HKU1)
R1AB_CVMJH
Murine coronavirus (strain JHM) (MHV-JHM) (Murine hepatitis virus)
R1AB_IBVBC
Avian infectious bronchitis virus (strain Beaudette CK) (IBV)
R1AB_IBVM
Avian infectious bronchitis virus (strain M41) (IBV)

17 entries

Probable RNA-directed RNA polymerase (EC 2.7.7.48) (Gag-Pol protein)

Select_allDeselect_all
RDRP_BYDVP
Barley yellow dwarf virus (isolate PAV) (BYDV) reference strain

Putative RNA-directed RNA polymerase (EC 2.7.7.48)

RDRP_CFMVN
Cocksfoot mottle virus (isolate Dactylis glomerata/Norway/CfMV-NO/1995) (CfMV) reference strain

Replicase polyprotein P2AB

RDRP_CRSVL
Carnation ringspot virus (isolate Lommel) (CRSV) reference strain

RNA-directed RNA polymerase (EC 2.7.7.48) (Protein p88)

RDRP_GLVWB
Giardia lamblia virus (isolate Wang) (GLV) reference strain
RDRP_GRVMC
Groundnut rosette virus (strain MC1) (GRV) reference strain

RNA-directed RNA polymerase (RdRp) (EC 2.7.7.48)

RDRP_PEMVW
Pea enation mosaic virus-1 (strain WSG) (PEMV-1) reference strain

Protein P1-P2

RDRP_PLRV1
Potato leafroll virus (strain Potato/Scotland/strain 1/1984) (PLrV) reference strain

Protein P1-P2

RDRP_PNLV
Poinsettia latent virus (isolate Euphorbia pulcherrima/Germany/Siepen/2005) (PnLV) (Poinsettia cryptic virus) reference strain

Protein P2-P3

RDRP_RCNMV
Red clover necrotic mosaic virus (RCNMV) reference strain

RNA-directed RNA polymerase (EC 2.7.7.48) (Protein p88)

RDRP_SBMVA
Southern bean mosaic virus (isolate Bean/United States/Arkansas) (SBMV) reference strain

Replicase polyprotein P2AB

RDRP_SCPMV
Southern cowpea mosaic virus (SCPMV) (Southern bean mosaic virus (strain cowpea)) reference strain

Replicase polyprotein P2AB

RDRP_SCVLA
Saccharomyces cerevisiae virus L-A (ScV-L-A) (ScVL1) reference strain
RDRP_TYYVF
Turnip yellows virus (isolate FL-1) (TuYV) (BWYV-FL1) reference strain

Protein P1-P2

RDRP_BYDV1
Barley yellow dwarf virus (isolate MAV-PS1) (BYDV)

Putative RNA-directed RNA polymerase (EC 2.7.7.48)

RDRP_BYDVR
Barley yellow dwarf virus (isolate P-PAV) (BYDV)

Putative RNA-directed RNA polymerase (EC 2.7.7.48)

RDRP_PLRVW
Potato leafroll virus (strain Potato/Netherlands/Wageningen/1989) (PLrV)

Protein P1-P2

RDRP_SCVLB
Saccharomyces cerevisiae virus L-BC (ScV-L-BC) (ScVLa)

9 entries

Replicase polyprotein 1ab (ORF1ab polyprotein)

Select_allDeselect_all
RPOA_EAVBU
Equine arteritis virus (strain Bucyrus) (EAV) reference strain
RPOA_PRRSB
Porcine reproductive and respiratory syndrome virus (strain HB-1) (PRRSV) reference strain
RPOA_PRRSL
Porcine reproductive and respiratory syndrome virus (strain Lelystad) (PRRSV) reference strain
RPOA_SHFV
Simian hemorrhagic fever virus (SHFV) reference strain
RPOA_LDVC
Lactate dehydrogenase elevating virus (strain C) (LDV)
RPOA_LDVP
Lactate dehydrogenase elevating virus (strain Plagemann) (LDV)
RPOA_PRRS1
Porcine reproductive and respiratory syndrome virus (strain 16244B) (PRRSV)
RPOA_PRRSR
Porcine reproductive and respiratory syndrome virus (strain VR-2332) (PRRSV)
RPOA_PRRSS
Porcine reproductive and respiratory syndrome virus (isolate Pig/United States/SD 01-08/2001) (PRRSV)

1 entry

Replicase polyprotein 1TF (ORF1ab polyprotein)

RPOTF_PRRSL
Porcine reproductive and respiratory syndrome virus (strain Lelystad) (PRRSV) reference strain

1 entry

Protein suppressor of silencing (Gene product 5.5)

SGS_BPT7
Enterobacteria phage T7 (Bacteriophage T7) reference strain

2 entries

Probable tail assembly protein gp41 (Gene product 41) (gp41) (Tail assembly chaperone)

Select_allDeselect_all
TAP_BPMU
Escherichia phage Mu (Bacteriophage Mu) reference strain
TAP_BPP2
Escherichia phage P2 (Bacteriophage P2) reference strain

Tail assembly protein E (Gene product E) (gpE) (Tail assembly chaperone) (TAC)

2 entries

Tail assembly protein Gp25 (Gene 25 protein) (Gp25)

Select_allDeselect_all
TAP25_BPMD2
Mycobacterium phage D29 (Mycobacteriophage D29) reference strain
TAP25_BPML5
Mycobacterium phage L5 (Mycobacteriophage L5) reference strain

2 entries

Probable tail assembly protein FS-gp41

Select_allDeselect_all
TAPFS_BPMU
Escherichia phage Mu (Bacteriophage Mu) reference strain
TAPFS_BPP2
Escherichia phage P2 (Bacteriophage P2) reference strain

Tail assembly protein E' (Gene product E') (gpE') (Tail assembly chaperone) (TAC)

1 entry

Tail tube protein gp17.1* (TTP) (Gene product 17.1) (gp17.1) (Major tail protein) (MTP)

TUBE_BPSPP
Bacillus phage SPP1 (Bacteriophage SPP1) reference strain

1 entry

Fusion protein 5.5/5.7

V5557_BPT7
Enterobacteria phage T7 (Bacteriophage T7) reference strain

2 entries

Tail assembly protein Gp24 (Gene product 24) (Gp24) (Minor tail protein Gp24)

Select_allDeselect_all
VG24_BPMD2
Mycobacterium phage D29 (Mycobacteriophage D29) reference strain
VG24_BPML5
Mycobacterium phage L5 (Mycobacteriophage L5) reference strain

1 entry

Bifunctional protease/dUTPase

VPRT_JSRV
Sheep pulmonary adenomatosis virus (Jaagsiekte sheep retrovirus) (JSRV)