Viral budding via the host ESCRT complexes


Many enveloped viruses recruit components of the cellular ESCRT (endosomal sorting complex required for transport) system to mediate host-assisted viral budding .

ESCRT complexes are normally used by the cell for biological functions involving membrane remodeling, such as intraluminal vesicle formation, autophagy or terminal stages of cytokinesis . The ESCRT family consists of ESCRT-0, ESCRT-I, ESCRT-II which are primarily involved in cargo sorting and membrane deformation, and ESCRT-III which cleaves the bud neck from its cytosolic face . In the last step, vps4 disassembles the complex. The budding reaction catalyzed by the ESCRT machinery has reversed topology when compared with most other budding processes in the cell, such as endocytosis and formation of transport vesicles.



Specific motifs in viral proteins called late assembly domains (L-domains) are responsible for recruiting the host ESCRT machinery directly or via ESCRT-associated proteins like ALIX.
- PTAP motif (and PSAP) recruits the ESCRT machinery via Tsg101.
- LYPxnL motif bind the adaptor protein ALIX, which in turn binds and recruits the ESCRT machinery.
- PPxY motif enlists members of the NEDD4 family of ubiquitin ligases through interactions with WW domains, although the precise link between the NEDD4-like proteins and the ESCRT machinery is still unknown.
- A fourth motif encoded by FPIV has been identified in the matrix protein of the paramyxovirus simian virus 5.
Some enveloped viruses however seem to use other mechanisms to bud, as it has been shown for Influenza virus .

Viruses budding through the ESCRT machinery:

Family Virus Viral protein L-domain motif Host ESCRT Budding location ref
Retroviridae HIV-1 P6-gag PTAP TSG101 Cell membrane
LYPxnL ALIX (low affinity) Cell membrane
SIV P6-gag PTAP TSG101 Cell membrane
LYPxnL ALIX Cell membrane
HTLV-1 P19-gag PPxY NEDD4 Cell membrane
PTAP TSG101 (low affinity) Cell membrane
Human spumaretrovirus Gag protein PTAP TSG101 Cell membrane
Moloney murine leukemia virus P12-gag PPxY ITCH Cell membrane
PTAP TSG101 Cell membrane
LYPxnL ALIX Cell membrane
Bovine leukemia virus Gag protein PPxYnL NEDD4 Cell membrane
Rous sarcoma virus Gag protein PPxY NEDD4 Cell membrane
Mason-Pfizer monkey virus Gag protein PSAP TSG101 Cell membrane
PPxY NEDD4 Cell membrane
EIAV P9-gag LYPxnL ALIX Cell membrane
Hepadnaviridae HBV ?Capsid protein ? ALIX Endoplasmic reticulum
Filoviridae Ebolavirus Matrix protein PTAP TSG101 Cell membrane
PPxY NEDD4 Cell membrane
Marburgvirus Matrix protein PPxY NEDD4 Cell membrane
Nucleoprotein PSAP TSG101 Cell membrane
Rhabdoviridae Vesicular stomatitis virus M protein PPxY NEDD4 Cell membrane
?PSAP ? Cell membrane
Rabies virus M protein PPxY ? Cell membrane
Paramyxoviridae Sendai virus M protein YLDL ALIX Cell membrane
Nipah virus M protein YMYL ? Cell membrane
Simian virus 5 M protein FPIV ? Cell membrane
Arenaviridae Lymphocytic choriomeningitis virus Z protein PTAP TSG101 Cell membrane
Lassa virus Z protein PTAP TSG101 Cell membrane
PPxY ? Cell membrane
Bunyaviridae Gn glycoproteins YxxL? ? Golgi
Flaviviridae Hepatitis C virus ?Core protein ? ? Endoplasmic reticulum
Herpesviridae Herpes simplex virus 1 ? ? Vps4 Trans-Golgi
Epstein-Barr virus ? ? ? Trans-Golgi
HCMV ? ? Vps4 Trans-Golgi
Poxviridae Vaccinia virus F13 protein LYPx(n)L ? Cell membrane

Matching UniProtKB/Swiss-Prot entries

207 entries grouped by strain (browse by keywords)

All proteins are shown (view only 137 complete proteome entries)

2 entries

Lake Victoria marburgvirus (strain Musoke-80) (MARV) (Marburg virus (strain Kenya/Musoke/1980)) reference strain

Select_all Deselect_all  
NCAP_MABVMNucleoprotein (Nucleocapsid protein) (Protein N)
VP40_MABVMMatrix protein VP40 (Membrane-associated protein VP40)

1 entry

Australian bat lyssavirus (isolate Bat/AUS/1996) (ABLV) reference strain


MATRX_ABLVBMatrix protein (Phosphoprotein M2)

1 entry

Chandipura virus (strain I653514) (CHPV) reference strain


MATRX_CHAVMatrix protein

1 entry

Drosophila melanogaster sigma virus (isolate Drosophila/USA/AP30/2005) (DMelSV) reference strain


MATRX_DMSVAMatrix protein

1 entry

European bat lyssavirus 1 (strain Bat/Germany/RV9/1968) (EBLV1) reference strain


MATRX_EBLV1Matrix protein

1 entry

European bat lyssavirus 2 (strain Human/Scotland/RV1333/2002) (EBLV2) reference strain


MATRX_EBLV2Matrix protein

1 entry

Human immunodeficiency virus type 1 group M subtype A (isolate MAL) (HIV-1) reference strain


GAG_HV1MAGag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 1 group M subtype B (isolate ARV2/SF2) (HIV-1) reference strain


GAG_HV1A2Gag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 1 group M subtype B (isolate BRU/LAI) (HIV-1) reference strain


GAG_HV1BRGag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 1 group M subtype B (isolate HXB2) (HIV-1) reference strain


GAG_HV1H2Gag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 1 group O (isolate ANT70) (HIV-1) reference strain


GAG_HV1ANGag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 2 subtype A (isolate BEN) (HIV-2) reference strain


GAG_HV2BEGag polyprotein (Pr55Gag)

1 entry

Human spumaretrovirus (SFVcpz(hu)) (Human foamy virus) reference strain


GAG_FOAMVGag polyprotein (Pr71Gag)

1 entry

Lassa virus (strain Mouse/Sierra Leone/Josiah/1976) (LASV) reference strain


Z_LASSJRING finger protein Z (Protein Z) (Zinc-binding protein)

1 entry

Latino mammarenavirus (isolate Rat/Bolivia/MARU 1924/1965) (LATV) reference strain


Z_LATVBRING finger protein Z (Protein Z) (Zinc-binding protein)

1 entry

Lymphocytic choriomeningitis virus (strain Armstrong) (LCMV) reference strain


Z_LYCVARING finger protein Z (Protein Z) (Zinc-binding protein)

1 entry

Mokola virus (MOKV) reference strain


MATRX_MOKVMatrix protein (Phosphoprotein M2)

1 entry

Moloney murine leukemia virus (isolate Shinnick) (MoMLV) reference strain


GAG_MLVMSGag polyprotein (Pr65gag) (Core polyprotein)

1 entry

Mouse mammary tumor virus (strain C3H) (MMTV) reference strain


GAG_MMTVCGag polyprotein

1 entry

Newcastle disease virus (strain Chicken/United States/B1/48) (NDV) reference strain


MATRX_NDVB1Matrix protein

1 entry

Parainfluenza virus 5 (strain W3) (PIV5) (Simian virus 5) reference strain


MATRX_PIV5Matrix protein (Membrane protein)

1 entry

Pirital mammarenavirus (isolate Rat/Venezuela/VAV-488/1995) (PIRV) reference strain


Z_PIRVVRING finger protein Z (Protein Z) (Zinc-binding protein)

1 entry

Rabies virus (strain Pasteur vaccins / PV) (RABV) reference strain


MATRX_RABVPMatrix protein (Phosphoprotein M2)

1 entry

Sendai virus (strain Ohita) (SeV) reference strain


MATRX_SENDOMatrix protein (M protein)

1 entry

Simian foamy virus (isolate chimpanzee) (SFVcpz) reference strain


GAG_SFVCPGag polyprotein (Pr71Gag)

1 entry

Simian foamy virus type 1 (SFVmac) (SFV-1) reference strain


GAG_SFV1Gag polyprotein (Pr71Gag)

1 entry

Simian immunodeficiency virus (isolate EK505) (SIV-cpz) (Chimpanzee immunodeficiency virus) reference strain


GAG_SIVEKGag polyprotein (Pr55Gag)

1 entry

Simian immunodeficiency virus (isolate MB66) (SIV-cpz) (Chimpanzee immunodeficiency virus) reference strain


GAG_SIVMBGag polyprotein (Pr55Gag)

1 entry

Simian immunodeficiency virus (isolate TAN1) (SIV-cpz) (Chimpanzee immunodeficiency virus) reference strain


GAG_SIVTNGag polyprotein (Pr55Gag)

1 entry

Tacaribe virus (strain Franze-Fernandez) (TCRV) reference strain


Z_TACVFRING finger protein Z (Protein Z) (Zinc-binding protein)

1 entry

Tamiami mammarenavirus (isolate Rat/United States/W 10777/1964) (TAMV) reference strain


Z_TAMVURING finger protein Z (Protein Z) (Zinc-binding protein)

1 entry

Tibrogargan virus (strain CS132) (TIBV) reference strain


MATRX_TIBVCMatrix protein

1 entry

Tupaia virus (isolate Tupaia/Thailand/-/1986) (TUPV) reference strain


MATRX_TUPVTMatrix protein

1 entry

Vaccinia virus (strain Western Reserve) (VACV) (Vaccinia virus (strain WR)) reference strain


F13_VACCWEnvelope protein F13 (37 kDa protein) (Palmitoylated EV membrane protein) (p37K)

1 entry

Variola virus (isolate Human/India/Ind3/1967) (VARV) (Smallpox virus) reference strain


F13_VAR67Envelope protein F13 (37 kDa protein) (Palmitoylated EV membrane protein) (p37K)

1 entry

Vesicular stomatitis Indiana virus (strain San Juan) (VSIV) reference strain


MATRX_VSIVAMatrix protein

1 entry

Vesicular stomatitis New Jersey virus (strain Ogden subtype Concan) (VSNJV) reference strain


MATRX_VSNJOMatrix protein

1 entry

Xenotropic MuLV-related virus (isolate VP62) (XMRV) reference strain


GAG_XMRV6Gag polyprotein (Pr65gag) (Core polyprotein)

1 entry

Zaire ebolavirus (strain Mayinga-76) (ZEBOV) (Zaire Ebola virus) reference strain


VP40_EBOZMMatrix protein VP40 (Membrane-associated protein VP40)

2 entries

Lake Victoria marburgvirus (strain Angola/2005) (MARV)

Select_all Deselect_all  
NCAP_MABVANucleoprotein (Nucleocapsid protein)
VP40_MABVAMatrix protein VP40 (Membrane-associated protein VP40)

2 entries

Lake Victoria marburgvirus (strain Ozolin-75) (MARV) (Marburg virus (strain South Africa/Ozolin/1975))

Select_all Deselect_all  
NCAP_MABVONucleoprotein (Nucleocapsid protein)
VP40_MABVOMatrix protein VP40 (Membrane-associated protein VP40)

2 entries

Lake Victoria marburgvirus (strain Popp-67) (MARV) (Marburg virus (strain West Germany/Popp/1967))

Select_all Deselect_all  
NCAP_MABVPNucleoprotein (Nucleocapsid protein) (Protein N)
VP40_MABVPMatrix protein VP40 (Membrane-associated protein VP40)

2 entries

Lake Victoria marburgvirus (strain Ravn-87) (MARV) (Marburg virus (strain Kenya/Ravn/1987))

Select_all Deselect_all  
NCAP_MABVRNucleoprotein (Nucleocapsid protein)
VP40_MABVRMatrix protein VP40 (Membrane-associated protein VP40)

1 entry

AKV murine leukemia virus (AKR (endogenous) murine leukemia virus)


GAG_MLVAVGag polyprotein (Core polyprotein)

1 entry

Abelson murine leukemia virus


GAG_MLVABGag polyprotein

1 entry

Allpahuayo mammarenavirus (isolate Rat/Peru/CLHP-2472/1997) (ALLV)


Z_ALLVPRING finger protein Z (Protein Z) (Zinc-binding protein)

1 entry

Aravan virus (ARAV)


MATRX_ARAVMatrix protein (Phosphoprotein M2)

1 entry

Australian bat lyssavirus (isolate Human/AUS/1998) (ABLV)


MATRX_ABLVHMatrix protein (Phosphoprotein M2)

1 entry

Avian myelocytomatosis virus HBI


GAG_AVIMDGag polyprotein

1 entry

Avian myelocytomatosis virus MC29


GAG_AVIMCGag polyprotein

1 entry

Avian sarcoma virus (strain CT10)


GAGC_AVISCP47(GAG-CRK) protein

1 entry

Avian sarcoma virus (strain Y73)


GAG_AVISYGag polyprotein

1 entry

Baboon endogenous virus (strain M7)


GAG_BAEVMGag polyprotein (Core polyprotein)

1 entry

Bear Canyon mammarenavirus (isolate Mouse/United States/AV A0070039/2000) (BCNV)


Z_BCNVURING finger protein Z (Protein Z) (Zinc-binding protein)

1 entry

Bovine immunodeficiency virus (strain R29) (BIV) (Bovine immunodeficiency-like virus)


GAG_BIV29Gag polyprotein (Pr53Gag)

1 entry

Bovine leukemia virus (isolate Australian) (BLV)


GAG_BLVAUGag polyprotein

1 entry

Bovine leukemia virus (isolate Japanese BLV-1) (BLV)


GAG_BLVJGag polyprotein

1 entry

Caprine arthritis encephalitis virus (strain Cork) (CAEV-Co)


GAG_CAEVCGag polyprotein

1 entry

Cas-Br-E murine leukemia virus


GAG_MLVCBGag polyprotein (Core polyprotein)

1 entry

Chapare mammarenavirus (isolate Human/Bolivia/810419/2003)


Z_CHAVBRING finger protein Z (Protein Z) (Zinc-binding protein)

1 entry

Cupixi mammarenavirus (isolate Rat/Brasil/BeAn 119303/1970) (CPXV)


Z_CPXVBRING finger protein Z (Protein Z) (Zinc-binding protein)

1 entry

Equine infectious anemia virus (isolate 1369) (EIAV)


GAG_EIAV9Gag polyprotein

1 entry

Equine infectious anemia virus (isolate CL22) (EIAV)


GAG_EIAVCGag polyprotein

1 entry

Equine infectious anemia virus (strain Wyoming) (EIAV)

1 entry

FBR murine osteosarcoma virus (FBR-MSV) (Finkel-Biskis-Reilly murine osteosarcoma virus)


GAG_MSVFRGag polyprotein (Core polyprotein)

1 entry

Feline immunodeficiency virus (isolate Petaluma) (FIV)


GAG_FIVPEGag polyprotein

1 entry

Feline immunodeficiency virus (isolate TM2) (FIV)


GAG_FIVT2Gag polyprotein

1 entry

Feline immunodeficiency virus (isolate Wo) (FIV)


GAG_FIVWOGag polyprotein

1 entry

Feline immunodeficiency virus (strain San Diego) (FIV)


GAG_FIVSDGag polyprotein

1 entry

Feline leukemia virus


GAG_FLVGag polyprotein (Core polyprotein)

1 entry

Feline sarcoma virus (strain Gardner-Arnstein) (Ga-FeSV) (Gardner-Arnstein feline leukemia oncovirus B)


GAG_FSVGAGag polyprotein (Core polyprotein)

1 entry

Feline sarcoma virus (strain McDonough)


GAG_FSVMDGag polyprotein (Core polyprotein)

1 entry

Feline sarcoma virus (strain Snyder-Theilen)


GAG_FSVSTGag polyprotein (Core polyprotein)

1 entry

Friend murine leukemia virus (isolate 57) (FrMLV)


GAG_MLVF5Gag polyprotein (Core polyprotein)

1 entry

Friend murine leukemia virus (isolate FB29) (FrMLV)


GAG_MLVFFGag polyprotein (Core polyprotein)

1 entry

Friend murine leukemia virus (isolate PVC-211) (FrMLV)


GAG_MLVFPGag polyprotein (Core polyprotein)

1 entry

Friend spleen focus-forming virus (isolate 502) (FSFFV)


GAG_FRSF5Gag polyprotein (Core polyprotein)

1 entry

Fujinami sarcoma virus


GAG_FUJSVGag polyprotein

1 entry

Gibbon ape leukemia virus (GALV)


GAG_GALVGag polyprotein (Core polyprotein)

1 entry

Guanarito mammarenavirus (isolate Human/Venezuela/NH-95551/1990) (GTOV)


Z_GTOVVRING finger protein Z (Protein Z) (Zinc-binding protein)

1 entry

Hortulanus murine leukemia virus (HoMuLV) (Mus hortulanus virus)


GAG_MLVHOGag polyprotein (Core polyprotein)

1 entry

Human immunodeficiency virus type 1 group M subtype A (isolate U455) (HIV-1)


GAG_HV1U4Gag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 1 group M subtype B (isolate BH10) (HIV-1)


GAG_HV1B1Gag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 1 group M subtype B (isolate BH5) (HIV-1)


GAG_HV1B5Gag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 1 group M subtype B (isolate CDC-451) (HIV-1)


GAG_HV1C4Gag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 1 group M subtype B (isolate JH32) (HIV-1)


GAG_HV1J3Gag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 1 group M subtype B (isolate JRCSF) (HIV-1)


GAG_HV1JRGag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 1 group M subtype B (isolate LW123) (HIV-1)


GAG_HV1LWGag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 1 group M subtype B (isolate MN) (HIV-1)


GAG_HV1MNGag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 1 group M subtype B (isolate NY5) (HIV-1)

1 entry

Human immunodeficiency virus type 1 group M subtype B (isolate OYI) (HIV-1)


GAG_HV1OYGag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 1 group M subtype B (isolate RF/HAT3) (HIV-1)


GAG_HV1RHGag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 1 group M subtype B (isolate YU-2) (HIV-1)


GAG_HV1Y2Gag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 1 group M subtype B (strain 89.6) (HIV-1)


GAG_HV1B9Gag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 1 group M subtype C (isolate 92BR025) (HIV-1)


GAG_HV192Gag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 1 group M subtype C (isolate ETH2220) (HIV-1)


GAG_HV1ETGag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 1 group M subtype D (isolate ELI) (HIV-1)


GAG_HV1ELGag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 1 group M subtype D (isolate NDK) (HIV-1)


GAG_HV1NDGag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 1 group M subtype D (isolate Z2/CDC-Z34) (HIV-1)


GAG_HV1Z2Gag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 1 group M subtype F1 (isolate 93BR020) (HIV-1)


GAG_HV193Gag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 1 group M subtype F1 (isolate VI850) (HIV-1)


GAG_HV1VIGag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 1 group M subtype F2 (isolate MP255) (HIV-1)


GAG_HV1MPGag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 1 group M subtype F2 (isolate MP257) (HIV-1)


GAG_HV1M2Gag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 1 group M subtype G (isolate 92NG083) (HIV-1)


GAG_HV19NGag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 1 group M subtype G (isolate SE6165) (HIV-1)


GAG_HV1SEGag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 1 group M subtype H (isolate 90CF056) (HIV-1)


GAG_HV190Gag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 1 group M subtype H (isolate VI991) (HIV-1)


GAG_HV1V9Gag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 1 group M subtype J (isolate SE9173) (HIV-1)


GAG_HV1S9Gag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 1 group M subtype J (isolate SE9280) (HIV-1)


GAG_HV1S2Gag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 1 group M subtype K (isolate 96CM-MP535) (HIV-1)


GAG_HV196Gag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 1 group M subtype K (isolate 97ZR-EQTB11) (HIV-1)


GAG_HV197Gag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 1 group N (isolate YBF106) (HIV-1)


GAG_HV1YBGag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 1 group N (isolate YBF30) (HIV-1)


GAG_HV1YFGag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 1 group O (isolate MVP5180) (HIV-1)


GAG_HV1MVGag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 2 subtype A (isolate CAM2) (HIV-2)


GAG_HV2CAGag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 2 subtype A (isolate D194) (HIV-2)


GAG_HV2D1Gag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 2 subtype A (isolate Ghana-1) (HIV-2)


GAG_HV2G1Gag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 2 subtype A (isolate KR) (HIV-2)


GAG_HV2KRGag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 2 subtype A (isolate NIH-Z) (HIV-2)


GAG_HV2NZGag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 2 subtype A (isolate ROD) (HIV-2)


GAG_HV2ROGag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 2 subtype A (isolate SBLISY) (HIV-2)


GAG_HV2SBGag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 2 subtype A (isolate ST) (HIV-2)


GAG_HV2STGag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 2 subtype B (isolate D205) (HIV-2)


GAG_HV2D2Gag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 2 subtype B (isolate EHO) (HIV-2)


GAG_HV2EHGag polyprotein (Pr55Gag)

1 entry

Human immunodeficiency virus type 2 subtype B (isolate UC1) (HIV-2)


GAG_HV2UCGag polyprotein (Pr55Gag)

1 entry

Ippy mammarenavirus (isolate Rat/Central African Republic/Dak An B 188 d/1970) (IPPYV)


Z_IPPYVRING finger protein Z (Protein Z) (Zinc-binding protein)

1 entry

Irkut virus (IRKV)


MATRX_IRKVMatrix protein (Phosphoprotein M2)

1 entry

Isfahan virus (ISFV)


MATRX_ISFVMatrix protein

1 entry

Jembrana disease virus (JDV)


GAG_JEMBRGag polyprotein (Pr53Gag)

1 entry

Junin mammarenavirus (JUNV) (Junn mammarenavirus)


Z_JUNINRING finger protein Z (Protein Z) (Zinc-binding protein)

1 entry

Khujand virus (KHUV)


MATRX_KHUVMatrix protein (Phosphoprotein M2)

1 entry

Koala retrovirus (KoRV)


GAG_KORVGag polyprotein (Core polyprotein)

1 entry

Lagos bat virus (LBV)


MATRX_LBVMatrix protein

1 entry

Machupo virus (MACV)


Z_MACHURING finger protein Z (Protein Z) (Zinc-binding protein)

1 entry

Maedi visna virus (strain 1514 / clone LV1-1KS1) (MVV) (Visna lentivirus)


GAG_VILV1Gag polyprotein

1 entry

Maedi visna virus (strain 1514 / clone LV1-1KS2) (MVV) (Visna lentivirus)


GAG_VILV2Gag polyprotein

1 entry

Maedi visna virus (strain 1514) (MVV) (Visna lentivirus)


GAG_VILVGag polyprotein

1 entry

Maedi visna virus (strain KV1772) (MVV) (Visna lentivirus)


GAG_VILVKGag polyprotein

1 entry

Mason-Pfizer monkey virus (MPMV) (Simian Mason-Pfizer virus)


GAG_MPMVGag polyprotein (Core polyprotein)

1 entry

Mobala mammarenavirus (isolate Rat/Central African Republic/Acar 3080/1983) (MOBV)


Z_MOBVCRING finger protein Z (Protein Z) (Zinc-binding protein)

1 entry

Moloney murine sarcoma virus (MoMSV)


GAG_MSVMOGag polyprotein (Core polyprotein)

1 entry

Moloney murine sarcoma virus (strain ts110) (MoMSV)


GAG_MSVMTGag polyprotein (Core polyprotein)

1 entry

Mouse mammary tumor virus (strain BR6) (MMTV)


GAG_MMTVBGag polyprotein

1 entry

Murine leukemia virus (strain BM5 eco)


GAG_MLVBMGag polyprotein (Core polyprotein)

1 entry

Newcastle disease virus (strain Beaudette C/45) (NDV)


MATRX_NDVBMatrix protein

1 entry

Newcastle disease virus (strain Chicken/Australia-Victoria/32) (NDV)


MATRX_NDVAMatrix protein

1 entry

Oliveros mammarenavirus (isolate Mouse/Argentina/RIID 3229/1990) (OLVV)


Z_OLVVARING finger protein Z (Protein Z) (Zinc-binding protein)

1 entry

Ovine maedi visna related virus (strain South Africa) (SA-OMVV) (Ovine lentivirus)


GAG_OMVVSGag polyprotein

1 entry

Parana mammarenavirus (isolate Rat/Paraguay/12056/1965) (PARV) (Paran mammarenavirus)


Z_PARVPRING finger protein Z (Protein Z) (Zinc-binding protein)

1 entry

Piry virus (PIRYV)


MATRX_PIRYVMatrix protein

1 entry

Rabies virus (strain CVS-11) (RABV)


MATRX_RABVCMatrix protein (Phosphoprotein M2)

1 entry

Rabies virus (strain China/DRV) (RABV)


MATRX_RABVDMatrix protein (Phosphoprotein M2)

1 entry

Rabies virus (strain China/MRV) (RABV)


MATRX_RABVRMatrix protein (Phosphoprotein M2)

1 entry

Rabies virus (strain ERA) (RABV)


MATRX_RABVEMatrix protein (Phosphoprotein M2)

1 entry

Rabies virus (strain HEP-Flury) (RABV)


MATRX_RABVHMatrix protein (Phosphoprotein M2)

1 entry

Rabies virus (strain India) (RABV)


MATRX_RABVIMatrix protein (Phosphoprotein M2)

1 entry

Rabies virus (strain Nishigahara RCEH) (RABV)


MATRX_RABVNMatrix protein (Phosphoprotein M2)

1 entry

Rabies virus (strain PM1503/AVO1) (RABV)


MATRX_RABVAMatrix protein (Phosphoprotein M2)

1 entry

Rabies virus (strain SAD B19) (RABV)


MATRX_RABVSMatrix protein (Phosphoprotein M2)

1 entry

Rabies virus (strain silver-haired bat-associated) (RABV) (SHBRV)


MATRX_RABVBMatrix protein (Phosphoprotein M2)

1 entry

Radiation murine leukemia virus


GAG_MLVRDGag polyprotein (Core polyprotein)

1 entry

Reston ebolavirus (strain Philippines-96) (REBOV) (Reston Ebola virus)


VP40_EBOREMatrix protein VP40 (Membrane-associated protein VP40)

1 entry

Reston ebolavirus (strain Reston-89) (REBOV) (Reston Ebola virus)


VP40_EBORRMatrix protein VP40 (Membrane-associated protein VP40)

1 entry

Sabia mammarenavirus (isolate Human/Brasil/SPH114202/1990) (SABV) (Sabi mammarenavirus)


Z_SABVBRING finger protein Z (Protein Z) (Zinc-binding protein)

1 entry

Sendai virus (strain Enders) (SeV)


MATRX_SENDEMatrix protein (M protein)

1 entry

Sendai virus (strain Fushimi) (SeV)


MATRX_SENDFMatrix protein (M protein)

1 entry

Sendai virus (strain Hamamatsu) (SeV)


MATRX_SENDAMatrix protein (M protein)

1 entry

Sendai virus (strain Harris) (SeV)


MATRX_SENDHMatrix protein (M protein)

1 entry

Sendai virus (strain Z) (SeV) (Sendai virus (strain HVJ))


MATRX_SENDZMatrix protein (M protein)

1 entry

Sheep pulmonary adenomatosis virus (Jaagsiekte sheep retrovirus) (JSRV)


GAG_JSRVGag polyprotein

1 entry

Simian foamy virus type 3 (strain LK3) (SFVagm) (SFV-3)


GAG_SFV3LGag polyprotein (Pr71Gag)

1 entry

Simian immunodeficiency virus (isolate CPZ GAB1) (SIV-cpz) (Chimpanzee immunodeficiency virus)


GAG_SIVCZGag polyprotein (Pr55Gag)

1 entry

Simian immunodeficiency virus (isolate F236/smH4) (SIV-sm) (Simian immunodeficiency virus sooty mangabey monkey)


GAG_SIVS4Gag polyprotein (Pr55Gag)

1 entry

Simian immunodeficiency virus (isolate GB1) (SIV-mnd) (Simian immunodeficiency virus mandrill)


GAG_SIVGBGag polyprotein (Pr55Gag)

1 entry

Simian immunodeficiency virus (isolate K6W) (SIV-mac) (Simian immunodeficiency virus rhesus monkey)


GAG_SIVMKGag polyprotein (Pr55Gag)

1 entry

Simian immunodeficiency virus (isolate Mm142-83) (SIV-mac) (Simian immunodeficiency virus rhesus monkey)


GAG_SIVM1Gag polyprotein (Pr55Gag)

1 entry

Simian immunodeficiency virus (isolate PBj14/BCL-3) (SIV-sm) (Simian immunodeficiency virus sooty mangabey monkey)


GAG_SIVSPGag polyprotein (Pr55Gag)

1 entry

Simian immunodeficiency virus (isolate STM) (SIV-mac) (Simian immunodeficiency virus rhesus monkey)


GAG_SIVMSGag polyprotein (Pr55Gag)

1 entry

Simian immunodeficiency virus agm.grivet (isolate AGM gr-1) (SIV-agm.gri) (Simian immunodeficiency virus African green monkey grivet)


GAG_SIVG1Gag polyprotein (Pr55Gag)

1 entry

Simian immunodeficiency virus agm.vervet (isolate AGM TYO-1) (SIV-agm.ver) (Simian immunodeficiency virus African green monkey vervet)


GAG_SIVVTGag polyprotein (Pr55Gag)

1 entry

Simian immunodeficiency virus agm.vervet (isolate AGM155) (SIV-agm.ver) (Simian immunodeficiency virus African green monkey vervet)


GAG_SIVV1Gag polyprotein (Pr55Gag)

1 entry

Simian immunodeficiency virus agm.vervet (isolate AGM3) (SIV-agm.ver) (Simian immunodeficiency virus African green monkey vervet)


GAG_SIVVGGag polyprotein (Pr55Gag)

1 entry

Simian retrovirus SRV-1


GAG_SRV1Gag polyprotein (Core polyprotein)

1 entry

Simian retrovirus SRV-2


GAG_SRV2Gag polyprotein (Core polyprotein)

1 entry

Spring viremia of carp virus (Rhabdovirus carpia)


MATRX_SVCVMatrix protein

1 entry

Squirrel monkey retrovirus (SMRV-H) (SMRV-HLB)


GAG_SMRVHGag polyprotein

1 entry

Sudan ebolavirus (strain Uganda-00) (SEBOV) (Sudan Ebola virus)


VP40_EBOSUMatrix protein VP40 (Membrane-associated protein VP40)

1 entry

Vaccinia virus (strain Ankara) (VACV)


F13_VACCAEnvelope protein F13 (37 kDa protein) (Palmitoylated EV membrane protein) (p37K)

1 entry

Vaccinia virus (strain Copenhagen) (VACV)


F13_VACCCEnvelope protein F13 (37 kDa protein) (Palmitoylated EV membrane protein) (p37K)

1 entry

Vaccinia virus (strain IHD-J) (VACV)


F13_VACCIEnvelope protein F13 (37 kDa protein) (Palmitoylated EV membrane protein) (p37K)

1 entry

Vaccinia virus (strain L-IVP) (VACV)


F13_VACCPEnvelope protein F13 (37 kDa protein) (Palmitoylated EV membrane protein) (p37K)

1 entry

Vesicular stomatitis Indiana virus (strain 85CLB South America) (VSIV)


MATRX_VSIVSMatrix protein

1 entry

Vesicular stomatitis Indiana virus (strain 94GUB Central America) (VSIV)


MATRX_VSIVCMatrix protein

1 entry

Vesicular stomatitis Indiana virus (strain 98COE North America) (VSIV)


MATRX_VSIVNMatrix protein

1 entry

Vesicular stomatitis Indiana virus (strain Glasgow) (VSIV)


MATRX_VSIVGMatrix protein

1 entry

West Caucasian bat virus (WCBV)


MATRX_WCBVMatrix protein (Phosphoprotein M2)

1 entry

Whitewater Arroyo mammarenavirus (isolate Rat/United States/AV 9310135/1995) (WWAV)


Z_WWAVURING finger protein Z (Protein Z) (Zinc-binding protein)

1 entry

Woolly monkey sarcoma virus (WMSV) (Smian sarcoma-associated virus)


GAG_WMSVGag polyprotein (Core polyprotein)

1 entry

Xenotropic MuLV-related virus (isolate VP35) (XMRV)


GAG_XMRV3Gag polyprotein (Pr65gag) (Core polyprotein)

1 entry

Xenotropic MuLV-related virus (isolate VP42) (XMRV)


GAG_XMRV4Gag polyprotein (Pr65gag) (Core polyprotein)

1 entry

Zaire ebolavirus (strain Gabon-94) (ZEBOV) (Zaire Ebola virus)


VP40_EBOG4Matrix protein VP40 (Membrane-associated protein VP40)

1 entry

Zaire ebolavirus (strain Kikwit-95) (ZEBOV) (Zaire Ebola virus)


VP40_EBOZ5Matrix protein VP40 (Membrane-associated protein VP40)