Varicosavirus (taxid:140295)

VIRION

Virions have rod-shaped particles with modal lengths of 320-360 nm and diameters of 18 nm. They do not have an envelope structure.

GENOME

The genome is segmented; bipartite. After deproteinization of purified virions, both single stranded and double stranded RNA are detected.

GENE EXPRESSION

These viruses contain a single coat protein (CP) with an Mr of approximately 48 kDa and do not possess an envelope structure.

ENZYMES

• RNA-dependent RNA polymerase [L]
• Mononega-type capping
  • RNA TPase, GTase, N7 Mtase, 2'O Mtase [L]

REPLICATION

CYTOPLASMIC

1. Virus penetrates into the cell by effraction or plasmodesmata transport.
2. Sequential transcription , viral mRNAs are capped and polyadenylated by polymerase stuttering in the cytoplasm.
3. Replication presumably starts when enough nucleoprotein is present to encapsidate neo-synthetized antigenomes and genomes.
4. The ribonucleocapsid can penetrate in neighboring plant cells through plasmodesmata transport