Double-Jelly roll fold, vertical
The double jelly-roll fold, vertical consists of two single jelly roll folds oriented vertically and connected by a short linker region. It is assembled in hexamers of T=25 to T=1200 capsids. Also called PRD1-adenovirus fold.
Origin:
Topology
The strands are traditionally labeled B through I for the historical reason that the first solved structure, the jelly-roll capsid protein of the tomato bushy stunt virus, had an additional strand A outside the common core of the fold. The sheets are composed of strands BIDG and CHEF, which are folded so that strand B is opposite strand C, I opposite H, and so on.(source wikipedia).


Cellular homologs of the double jelly-roll major capsid proteins clarify the origins of an ancient virus kingdom
Mart Krupovic a, Kira S Makarova b, Eugene V Koonin b
Proc Natl Acad Sci U S A. 2022 Jan 25;119(5):e2120620119.
The logic of virus evolution
Eugene V Koonin, Valerian V Dolja, Mart Krupovic
Cell Host Microbe. 2022 Jul 13;30(7):917-929.
poxvirus-> D13 protein for immature PMID: 35361762